Lycoriella excavata, Menzel & Vilkamaa, 2021

Menzel, Frank & Vilkamaa, Pekka, 2021, New species and records of Lycoriella Frey (Diptera, Sciaridae) from the Holarctic region, Zootaxa 5072 (6), pp. 501-530 : 509-510

publication ID	https://doi.org/ 10.11646/zootaxa.5072.6.1
publication LSID	lsid:zoobank.org:pub:B1A0C8F3-692F-422E-8F20-35CC389DDD0E
DOI	https://doi.org/10.5281/zenodo.5751539
persistent identifier	https://treatment.plazi.org/id/03E1D765-D546-5105-FF72-FA510D380BF4
treatment provided by	Plazi
scientific name	Lycoriella excavata
status	sp. nov.

Treatment

Lycoriella excavata View in CoL sp. n.

Figs 2B View FIGURE 2 , 3D View FIGURE 3 , 12B View FIGURE 12

Material studied. Holotype male. USA, Colorado, Laramer Co., Pinewood Springs , 40.2677°N, 105.3678°W, Malaise trap, 26.VII–7.IX.2014, S. & B. Fitzgerald (in MZH, http://id.luomus.fi/GE.1811). GoogleMaps

Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 1–2 facets wide. Face with 27 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3 setae, with deep dorsal pit with sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.8x as long as wide, neck slightly longer than wide, longest setae longer than width of flagellomere. Thorax. Brown; setae pale. Anterior pronotum with 9 setae. Proepisternum with 10 setae. Scutellum with 2 moderately long and some short and fine setae. Wing. Fumose. Length 1.9 mm. Width/length 0.40. Anal lobe small. Veins distinct, except for stM. c/w 0.65. R1/R 0.75. stM longer than fork of M. bM as long as r-m, stCuA shortest. bM and r-m non-setose. Halter yellow. Legs. Pale brown; setae pale. Fore tibial organ forming large patch of setae in demarcated depression. Fore tibial spur longer than apical width of tibia.

Abdomen. Pale brown; setae pale, short and weak. Hypopygium ( Fig. 2B View FIGURE 2 ). Brown, like abdomen. Intergonocoxal area moderately long, with semicircular setose lobe. Gonocoxa narrow and long, longer than gonostylus, with sparse, rather short setosity. Gonostylus ( Fig. 3D View FIGURE 3 ) widest subapically, slightly narrowed towards apex, deeply excavated dorsally; densely setose apically, with long and narrow apical tooth, with 1 subapical and 7–8 medial megasetae on ventral margin of apical half of gonostylus; with long whip-lash seta in middle. Tegmen ( Fig. 12B View FIGURE 12 ) longer than wide, slightly notched apically, weakly sclerotized laterally, with area of minute teeth apicoventrally, with area of small aedeagal teeth. Aedeagal apodeme moderately long.

BIN. Unknown.

Discussion. Lycoriella excavata sp. n. with its subapically widened, lobe-shaped gonostyli resembles Lycoriella piristylata Vilkamaa, Hippa & Heller, 2013 , which has (in contrast to the original description) also a palpal sensory pit and therefore belongs to Lycoriella Frey. In contrast to Lycoriella excavata sp. n., in L. piristylata a long whiplash seta is missing, the intercoxal lobe is small (or absent in some specimens), the apical tooth is significantly shorter or completely reduced and the tegmen is short and trapezoidal ( Vilkamaa et al. 2013: fig. 3A–C). Lycoriella excavata sp. n. ( Fig. 2B View FIGURE 2 ) also differs from this species in its ventrally enlarged lobe-shaped gonostylus with a very deep dorsal excavation, all medial megasetae of the gonostylus on its ventral margin, and its tegmen with apicolateral teeth. Apicolateral teeth of the tegmen are currently known only in Lycoriella taimyrensis described here ( Fig. 13E View FIGURE 13 ), in some species of angustostylata species group in Bradysia Winnertz, 1867 ( Menzel & Heller 2005; Mohrig 2016), as well in Odontosciara Rübsaamen, 1908 ( Mohrig 2003) .