Gonaphodioides columbicus ( Harold, 1880 ) Dellacasa & Dellacasa & Gordon, 2012

Dellacasa, Marco, Dellacasa, Giovanni & Gordon, Robert D., 2012, Systematic revision of Gonaphodiellus taxa, with description of two new genera and fourteen new species (Coleoptera: Scarabaeidae: Aphodiinae), Insecta Mundi 2012 (230), pp. 1-41 : 18-19

Gonaphodioides columbicus ( Harold, 1880) new combination

( Fig. 78-83 )

Aphodius columbicus Harold, 1880: 36 ; Blackwelder 1944: 212.

Aphodius (Gonaphodiellu s) columbicus ; Schmidt 1913: 134; 1922: 121; Dellacasa 1988: 314.

Gonaphodiellus (s. l.) columbicus ; Skelley et al. 2007: 4 .

Type locality. Bogotá [ Colombia].

Type repository. Muséum national d’Histoire naturelle, Paris, France (lectotype herein designated).

Redescription. Length 4.0-5.0 mm, elongate, moderately convex, shiny, glabrous. Chestnut-brown; clypeal margin, sides of pronotum, apex of elytra shadowy reddish; legs pale reddish; antennal club testaceous. Head with epistome feebly gibbous, finely regularly and rather sparsely punctured throughout; clypeus slightly sinuate at middle, obtusely angulate at sides, faintly bordered; edge distinct and somewhat upturned anteriorly only, glabrous; genae subangulose, not ciliate, protruding from the eyes; frontal suture finely impressed; front finely and sparsely punctured. Pronotum transverse, moderately convex, finely, regularly and rather sparsely punctured; punctation somewhat denser anteriorly; sides feebly curved, very finely bordered, edge glabrous; hind angles widely truncate, truncature slightly inwardly sinuate; base distinctly bisinuate. Scutellum ogival elongate, somewhat depressed and distinctly microreticulate basally. Elytra convex, elongate with humeral denticle very small; striae fine, superficially punctured, not crenulate; interstriae distinctly convex on disc and nearly imperceptibly punctured laterally, nearly flattened subapically. Hind tibiae upper spur as long as half of first tarsal segment; first tarsal segment longer than following three segments combined. Male: pronotum relatively more transverse and less convex, parallel-sided with anterior discal punctation finer; elytra more elongate, nearly parallel-sided; fore tibiae more slender and more elongate, their spur downward bent; aedeagus Fig. 82-83 . Female: pronotum relatively less transverse and more convex, somewhat narrowed anteriorly, uniformly punctured throughout; elytra shorter and somewhat broadened backward; fore tibiae more stout and shorter, their spur straight.

Type material. Lectotype, male, herein designated ( MNHN), bearing the following labels: 1) white, handwritten: Bogotá; 2) white, printed: ex Musaeo/E. Steinheil; 3) red, printed: Aphodius / columbicus / Harold, 1880; 4) red, printed: Lectotypus/ M. & G. Dellacasa des. 2001 .

Paralectotypes: 3 males and 4 females from Bogotá (paralectotypes, MNHN) ; 3 females from La Luzera [ Colombia], ex Musaeo / E. Steinheil (paralectotypes, MNHN) ; 1 male and 1 female from Viani, ex Musaeo / Harold (paralectotypes, , MNHN) ; 1 female from Noescen. [ Colombia?] (paralectotype, MNHN) ; 1 female from Colombia, ex Musaeo / E. Harold (paralectotype, MNHN) .

Material examined. COLOMBIA: Agnas Largas près Bogotá, 1877, leg. Thieme O. (1 ex., MNHN) ; Bogotá (1 ex., DCGI) ; Bogotá, 1877, leg. Thieme O. (3 exx., MNHN) ; Oberhalb Bogotá, 1000’, 1877, leg. Thieme O. (3 exx., MNHN) ; Papayan , 1899, leg. Abbé Gaujon (1 ex., MNHN) ; Santander, R. Natural Cachalú, m 2150-2250, III.1999, leg. Gonzales F. (2 exx., DCGI) . VENEZUELA: Merida, Mucuy Fish Hatchery , 7 Km E Tabay, 6600’, 10-13.II.1978, leg. Heppner J. B., black light (3 exx., DCGI) ; Merida, La Mucuy Estacion , 10-15.VIII.1976, leg. Bordon C. (4 exx., DCGI) ; Merida, La Mucuy , m 2300, 10.IX.1965, leg. Bordon C. (1 ex., DCGI) ; Trujillo, Paramo del Guaramacal , m 3000, 24.V.1980, leg. Bordon C. (5 exx., DCGI) .

Distribution. Colombia, Venezuela.

Bionomics. Poorly known; the specimens examined were collected in spring, summer and early autumn.

Figures

Figure 78-88. Gonaphodioides columbicus (Harold, 1880) (Oberhalb Bogotá, Colombia). 78) Epipharynx. 79) Right labial palp (dorsal view). 80-81) Habitus (morphological details and length ideogram). 82-83) Aedeagus (dorsal and lateral view). Gonaphodioides newtoni, new species (3 mi. N Tlanchinol, Hidalgo, Mexico). 84) Epipharynx. 85-86) Habitus (morphological details and length ideogram). 87-88) Aedeagus (dorsal and lateral view).