Scarabaeidae, Latreille, 1802

Krell, Frank- Thorsten, 2006, Fossil Record And Evolution Of Scarabaeoidea (Coleoptera: Polyphaga), The Coleopterists Bulletin (mo 5) 60, pp. 120-143: 127-128

publication ID

http://doi.org/ 10.1649/0010-065x(2006)60[120:fraeos]2.0.co;2

persistent identifier

http://treatment.plazi.org/id/03E287E2-FFFB-FFDF-57F8-06C96244FD6C

treatment provided by

Carolina

scientific name

Scarabaeidae
status

 

Scarabaeidae  : PROTOTROGINAE Nikolajev 2000 a  (L Cretaceous)

Diagnosis: Mesocoxae elongate, contiguous; mesotibia with two transverse keels; elytra covering pygidium; five visible sternites; lateral margins of sternites forming sharp edges. Remarks: According to Nikolajev, the taxon is diagnosed by symplesiomorphies only (in respect to Trogidae  ). Since Nikolajev classifies Trogidae  , Geotrupidae  , and Scarabaeidae  as subfamilies (of Scarabaeidae  5 Scarabaeoidea  without Lucanidae  and Passalidae  ), his Prototroginae  should be upgraded to family level to be consistent with the current classification ( Lawrence et al. 2000; Scholtz and Gebennikov 2005). However, since Prototroginae  are diagnosed solely by symplesiomorphies ( Nikolajev 2000 a) and are likely to be paraphyletic, I refrain from formally giving it the new status of a family. Nevertheless, there are no extant scarabaeoids with five visible sternites and tibiae with two transverse keels.

Scarabaeidae  : Melolonthinae  : CRETOMELOLONTHINI Nikolajev 1998 (L Cretaceous) Diagnosis: Labrum not covered by clypeus; clypeus without anterior border; anterior border of pronotum leathery; Radius 1 (5 RA3) and R3 (5 RA4) apically approximated (but not fused); meso- and metatibia with single transverse carina; apical spurs of meso- and metatibia close-set; pygidium free; two last abdominal spiracles on sternites; six visible sternites. Remarks: All diagnostic characters seem to be plesiomorphic with respect to Melolonthinae  (wing venation) or even Scarabaeidae  . Cretomelolonthinae might be paraphyletic as defined.

Scarabaeidae  : Aclopinae  ?: HOLCOROBEINI Nikolajev 1992  (U Jurassic–L Cretaceous) Diagnosis: Medium-sized body; mandibles and labrum exposed; outer margin of protibia with three denticles; meso- and metatibia with two transverse keels; mesocoxae contiguous; each elytron with ten ‘dot-like grooves’; radius gradually thinning toward apex and running along frontal margin of hind wing; radius sector (5 RA4) curving smoothly without joining radius (5 RA3). Remarks: Nikolajev (2004) integrated the Holcorobeini  into Aclopinae  although two characters do not fit his own diagnosis of this subfamily: the tarsi of Holcorobeini  are not or only slightly longer than the tibia (much longer in Aclopinae  with the exception of Xenaclopus Arrow  ), and the meso- and metatibiae can have more than one keel on the outer side (one keel in Aclopinae  with a rudimentary second in Xenaclopus  ). Moreover, the metaventrite and coxae are much smaller in the fossils than in extant Aclopinae  .

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Loc

Scarabaeidae

Krell, Frank- Thorsten 2006
2006
Loc

HOLCOROBEINI

Nikolajev 1992
1992
Loc

Holcorobeini

Nikolajev 1992
1992
Loc

Holcorobeini

Nikolajev 1992
1992
Loc

Xenaclopus

Arrow 1915
1915
Loc

Xenaclopus

Arrow 1915
1915
Loc

Scarabaeidae

Latreille 1802
1802