Plusioglyphiulus saksit, Golovatch & Geoffroy & Mauriès & Vandenspiegel, 2011

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Vandenspiegel, Didier, 2011, The millipede genus Plusioglyphiulus Silvestri, 1923 in Thailand (Diplopoda, Spirostreptida, Cambalopsidae) 2940, Zootaxa 2940 (1), pp. 1-63 : 46-49

Plusioglyphiulus saksit sp. n. Figs 42−44 .

Material examined: Holotype male ( MNHN GA 068 ), Thailand, Chaiyaphum Prov., Khon San Distr., Cave Tham Ngoem 2, hand collection, 17 June 1987, leg. L. Deharveng & A. Bedos (THA-CYM-029) . Paratype: 1 female ( SEM), same locality, together with holotype .

Name: To emphasize “saksit”, in Thai meaning “holy”, a noun in apposition.

Diagnosis: Differs from congeners in the oligotrichous gnathochilarium, coupled with the shorter accessory filament at the claw base, the particular carinotaxy patterns, and the special shapes and armature of both gonopod pairs (see also Key below).

Description: Coloration uniformly grey-brown; head, antennae, collum, venter and legs light yellowish to pallid; metatergal crests and, especially, poriferous tubercles infuscate, dark brown.

Length of holotype about 38 mm, width 1.8 mm, with 62p+1a+T. Collum broadest.

All characters as in P. panhai sp. n. ( Figs 42E, F , 43C–E, G , 44A, E, F ), except as follows. Ocellaria small, dark brown, ovoid in shape, with about eight visible ocelli arranged in 4–5 longitudinal rows. Gnathochilarium ( Fig. 44B ) rather oligotrichous (n=2).

Postcollum constriction very evident, due to especially enlarged collum and segment 2 ( Figs 42A, D , 43A ). Carinotaxy formula of collum: /(t)/+1p/t+2p/t+3p/t+4p/t+/t/t+pp/t/t+/ma/m ( Figs 42A, D , 43B ). Carinotaxy of metaterga 2–4, 7/7+m/m+7/7; usual formula of all following metaterga, 3/3+I/i+3/3/3+m/m/m ( Figs 42A–C , 43F ); all crests and tubercles, including poriferous cones, rather low. Midbody segments ovoid in cross-section, slightly compressed laterally ( Fig. 43F ).

Male legs 1 with a usual strong central hook curved forward, appendages large, sac-shaped, densely setose, 1- segmented ( Fig. 44C ). Male legs 2 strongly enlarged, penes bare ( Fig. 44D ).

Anterior gonopods rather simple, with a paramedian pair of anterior coxosternal processes (cxp1) rather high, slender, only parabasally subcontiguous, suberect, apically forming a twisted subtriangular expansion; caudal pair of coxosternal processes (cxp2) only slightly higher, also suberect, each rounded and laterally supplied with a sharp terminal tooth on caudal face; telopodites (te) coniform, long, apically setose, about as high as cxp2, attached to coxal region caudolaterally, probably capable of movement ( Fig. 44G ). Posterior gonopods somewhat shorter, rather simple, coxites well separated from sternum, fused only basally, membranous and sac-shaped; each coxite mainly consisting of a stalk surmounted by a large fovea and a distinct flagellum; telopodites missing ( Fig. 44H ).

Figures

FIGURE 42. Plusioglyphiulus saksit sp. n., female paratype. A: anterior part of body, lateral view. B: midbody segments, lateral view. C: posterior part of body, lateral view. D: anterior part of body, dorsal view. E: midbody segments, dorsal view. F: posterior part of body, dorsal view. Scale bars: A–E, 0.5 mm; F, 0.2 mm.

FIGURE 43. Plusioglyphiulus saksit sp. n., female paratype. A: anterior part of body, ventral view. B: same, front view. C: midbody segments, ventral view. D: posterior part of body, ventral view. E: same, caudal view. F: cross-section of a midbody segment, caudal view. G: enlarged claw, sublateral view. Scale bars: A–D and F, 0.5 mm; E, 0.2 mm; G, 0.02 mm.

FIGURE 44. Plusioglyphiulus saksit sp. n., holotype. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, lateral view. D: leg 2, caudal view. E: leg 3, caudal view. F: midbody leg, lateral view. G: anterior gonopods, front view. H: posterior gonopods, caudal view. te: telopodite; cxp1: anterior coxosternal process; cxp2: posterior coxosternal process. Scale bar: A–F, 0.4 mm; G and H, 0.2 mm.