Enghophyllum sifnium, Lazányi & Vagalinski, 2013

Lazányi, Eszter & Vagalinski, Boyan, 2013, A new genus of the millipede tribe Brachyiulini (Diplopoda: Julida: Julidae) from the Aegean region, European Journal of Taxonomy 70, pp. 1-12 : 4-6

Enghophyllum sifnium gen. et sp. nov.

urn:lsid:zoobank.org:act:5327BB09-7438-47BD-9D91-BBC6FE7F1CC5

Figs 1 A-D, F, H-I, L-M, 2A-G, 5

Diagnosis

Differs from the only congener, E. naxium (Verhoeff, 1903) , by the apical margin of the promere being slightly convex and the opishtomere being significantly compressed meso-laterad.

Etymology

Named after the type locality, the island of Sifnos, Cyclades.

Material examined

Holotype

GREECE: ♂, Hellas [Greece], Kikladhes [Cyclades], Sifnos Island, Pharos [Faros], 11.ii.1990 [11 Feb. 1990], S. Sfendourakis leg., ex coll. Karamaouna, “ Megaphyllum aff. naxium ( Verhoeff, 1901) ” M. Karamouna det. ( ZMUC).

Paratypes

GREECE: 1 juv. ♂, 1 juv. ♀. Same collection data as holotype ( ZMUC).

Description

Holotype male in stadium X (with 9 rows of ocelli); body ring number: 41 podous + 2 apodous + telson; length 18.8 mm; height 1.1 mm; ocelli: right side: 35, left side: 33. Additional measurements of the adult male: promentum around twice as long as wide ( Fig. 1B ), tibia of midbody legs slightly longer than tarsus, 4 th antennal article as long as 5 th ( Fig. 1A ). Subadult paratype male in stadium IX (with 8 rows of ocelli); body ring number: 38 podous + 3 apodous + telson; length 13.1 mm; height 1.1 mm; ocelli: right side: 34, left side: 35. Subadult female in stadium IX (with 8 rows of ocelli); body ring number: 39 podous + 3 apodous + telson; length 14.1 mm; height 1 mm; ocelli: right side: 30, left side: 30. 17-18 labral bristles.

COLOUR. Basic colour yellowish-beige (obviously faded) with a narrow brownish middorsal band, divided by a white median line. Two wider, diffused brownish bands at ozopore level, more intensely coloured around the ozopores.

SCULPTURE. Collum smooth, with only several short striae visible; metazona sparsely striated ( Fig. 1C ), somewhat denser on dorsal side, posterior margin with long, erect setae. Scarce, short striae also on prozona.

TELSON ( Fig. 1D ). Anal valves covered with scarce, long setae and a row of much shorter setae along each valve’s caudal margin. Preanal process without any setae in the adult male, but with a few setae in the two subadults; straight, stout, protruding near to the level of the longest anal setae (for the adult male). Subanal scale triangular, with a rounded tip, slightly protruding behind rear contour of anal valves; with two pairs of long setae laterally.

GONOPODS ( Fig. 2 A-G). Promere ( Fig. 2 D-E and P on Fig. 2 A-C) wide, slightly narrowing apically towards mesal margin. Apical margin convex, mesal and lateral margins almost parallel, the lateral one slightly sinuous. Opisthomere with three processes: a flat lateral process (lp on Fig. 2 A-C, F), a slightly curved, rod-like basal posterior process (bpp on Fig. 2 A-C, F-G) and an elongate, massive, rod-like apical posterior process (app on Fig. 2 A-C, F-G); all processes well-developed, tapering and not furcated. A micro-spinose pillow (msp) around the base of solenomere (s) ( Fig. 2 A-C, F-G). Entire opisthomere strongly compressed meso-laterad (compare Fig. 2C with Fig. 4A ).

Female adults unknown.

Figures

Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet.

Fig. 2. Gonopods of Enghophyllum sifnium gen. et sp. nov., holotype ♂. A. Left gonopods, mesal view. B. Leftgonopods,antero-lateralview.C.Leftgonopods,meso-posteriorview.D.Leftpromere,meso-posterior view. E. Left promere, postero-mesal view. F. Right opisthomere, lateral view. G. Right opisthomere, mesal view. – app = apical posterior process; bpp = basal posterior process; fc = flagellum channel; fl = flagellum; fsi = flagellum sinus; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; tl = triangular lamella. Scale bar: 0.2 mm.

Fig. 4. Gonopods of Enghophyllum naxium, male from Mavri Islet. A. Gonopods, posterior view. B. Gonopods, antero-lateral view. C. Right gonopods, mesal view. D. Left opisthomere, lateral view. – app = apical posterior process; bpp = basal posterior process; d = deepening; fc = flagellum channel; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; sc = sperm canal; tl = triangular lamella. Scale bar: 0.2 mm.