Oxytelus subtilis, Sharp, 1889

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1067-1806 : 1067-1806

publication ID	https://doi.org/ 10.1206/0003-0090.265.1.1
persistent identifier	https://treatment.plazi.org/id/03E64505-FF8E-5F2A-FF0E-A287FE2796BA
treatment provided by	Felipe
scientific name	Oxytelus subtilis
status

Treatment

subtilis Sharp, 1889 View in CoL , see: humilis Erichson, 1840 .

sulciceps Bernhauer, 1926d: 247 ( Eleusis ; Type locality: Luzon, Mount Banahao; Provinz Nueva Vizcaya, Imugan ).

— Scheerpeltz, 1933: 999 ( Eleusis ; catalog).

- DISTRIBUTION: Philippines.

sulcicollis Scheerpeltz, 1956a: 480 , 493, 499 ( Eleusis ; Type locality: Belgisch-Kongo : Hembe Bitale. Also cited from French Congo).

- DISTRIBUTION: French Equatorial Africa, Congo.

suturalis Cameron, 1940c: 210 ( Eleusis ; Type locality: Ghum District : Tiger Hills, alt. 8,500–10,000 feet).

- DISTRIBUTION: India.

teestaensis Herman, 2001: 39 ( Eleusis ; replacement name for sikkimensis Scheerpeltz ).

- DISTRIBUTION: India.

sikkimensis Scheerpeltz, 1965: 128 ( Eleusis ; [preoccupied]; Type locality: British Sikkim, 200 m, Valley at Tista Bridge).

— Herman, 2001: 39 ( Eleusis ; synonym of teestaensis ).

tenuis Fauvel, 1865 , see: humilis Erichson, 1840 .

tenuis Walker, 1859 , see: humilis Erichson, 1840 .

terminata Fauvel, 1869: 494 ( Eleusis ; replacement name for apicipennis Kraatz ).

— Bernhauer, 1926d: 251 ( Eleusis ; characters in key).

— Herman, 2001: 22 ( Eleusis ; synonymic notes).

- DISTRIBUTION: Seychelles, Sri Lanka, Malaysia, Singapore, Indonesia, Taiwan, Japan, China.

apicipennis Kraatz, 1859: 183 ( Isomalus ; [preoccupied]; Type locality: Ceylan).

— Bernhauer and Schubert, 1910: 11 ( Eleusis ; synonym of kraatzi ).

— Bernhauer, 1922b: 165 ( Eleusis ; synonym of kraatzi ).

— Cameron, 1930: 85 ( Eleusis ; synonym of kraatzi ).

— Scheerpeltz, 1957c: 222 ( Eleusis ; synonym of kraatzi ).

kraatzi Fauvel, 1878f: 207 ( Eleusis ; replacement name for apicipennis Kraatz ).

— Bernhauer, 1902b: 45 ( Eleusis ; Ceylon).

— Fauvel, 1905e: 78 ( Eleusis ; Java; Celebes; Lombok; Engano Island; Nias Island; Sumatra; China; Ceylon; Woodlark Island).

— Bernhauer and Schubert, 1910: 11 ( Eleusis ; catalog).

— Cameron, 1921a: 362, 398 ( Eleusis ; characters; Singapore).

— Bernhauer, 1922a: 149 ( Eleusis ; Indochina).

— Bernhauer, 1922b: 165 ( Eleusis ; Seychelles).

— Bernhauer, 1922c: 221 ( Eleusis ; Formosa).

— Cameron, 1928d: 92 ( Eleusis ; illustrations; Sumatra).

— Bernhauer, 1928: 1 ( Eleusis ; Sumatra).

— Cameron, 1930: 85 ( Eleusis ; characters; Ceylon; Malay Peninsula; Singapore; Java).

— Scheerpeltz, 1933: 997 ( Eleusis ; catalog).

— Cameron, 1936a: 25 ( Eleusis ; Java).

— Cameron, 1949b: 456 ( Eleusis ; Formosa).

— Scheerpeltz, 1957c: 222 ( Eleusis ; Sumba).

— Jarrige, 1963a: 120 ( Eleusis ; Ceylon).

— Frank, 1982: 6 ( Eleusis ; list of parasites).

— Naomi, 1996: 49, 51 ( Eleusis ; characters; Japan).

— Herman, 2001: 22 ( Eleusis ; synonym of terminata ).

testacea Erichson, 1840: 840 ( Isomalus ; Type locality: Madagascar).

— Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Scheerpeltz, 1961g: 244 ( Eleusis ; characters; Madagascar).

tibialis Laporte, 1835: 131 ( Eleusis ; Type locality: Madagascar).

— Laporte, 1840: 186 ( Eleusis ; characters; Madagascar).

— Fauvel, 1865: 36 ( Isomalus ; characters; Madagascar).

— Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Scheerpeltz, 1961g: 241 ( Eleusis ; characters; Madagascar).

- DISTRIBUTION: Madagascar.

tonkinensis Cameron, 1939a: 22 ( Eleusis ; Type locality: Tonkin: Hoa Binh).

- DISTRIBUTION: Vietnam.

toxopei Cameron, 1952: 241 ( Eleusis ; Type locality: New Guinea, Snow Mountains , Moss Forest Camp, 2600 m).

- DISTRIBUTION: New Guinea.

transversiceps Scheerpeltz, 1935a: 595 ( Eleusis ; Type locality: Tjisaroea , Java).

- DISTRIBUTION: Indonesia.

turneri Bernhauer, 1940: 129 ( Eleusis ; Type locality: Kenya: Mt. Kinangop , bamboo zone).

- DISTRIBUTION: Kenya.

unicolor Cameron, 1942b: 321 ( Eleusis ; Type locality: Chyulu Hills, altitude 5600 feet).

- DISTRIBUTION: Kenya.

usambarae Bernhauer, 1912f: 177 ( Eleusis ; Type locality: Usambara).

— Scheerpeltz, 1933: 999 ( Eleusis ; catalog).

— Scheerpeltz, 1956a: 489 ( Eleusis ; characters; Usambara).

- DISTRIBUTION: Tanzania.

ustulatipennis Perroud and Montrousier, 1864 [Note: Described in Isomalus and transferred to “Clavicornes Ino ?” by Fauvel, 1869: 494], see: Inopeplidae .

validicornis Scheerpeltz, 1956a: 481 , 496, 505 ( Eleusis ; Type locality: Belgisch-Kongo : Hembe Bitale. Also cited from French Congo).

— Scheerpeltz, 1956a: 496 ( Eleusis ; characters; French Congo; Belgian Congo).

- DISTRIBUTION: French Equatorial Africa, Congo.

vermicollis Bernhauer, 1929b: 81 ( Eleusis ; Type locality: Haut Uele: Watsa).

— Scheerpeltz, 1933: 999 ( Eleusis ; catalog).

— Scheerpeltz, 1956a: 490 ( Eleusis ; characters; French Congo; Belgian Congo).

- DISTRIBUTION: French Equatorial Africa, Congo.

vermicularis Fauvel, 1895: 189 ( Eleusis ; Type locality: Bírmanie, Carin Cheba, 900–1100 m). — Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Cameron, 1930: 77 ( Eleusis ; characters; Burma).

— Scheerpeltz, 1933: 999 ( Eleusis ; catalog).

- DISTRIBUTION: Myanmar.

virgula Fauvel, 1889 , see: Zeoleusis .

viridans Fauvel, 1895: 189 ( Eleusis ; Type locality: Bírmanie, Carin Cheba , 900–1100 m; Carin Asciuii Cheba , 1200–1300 m; Tenasserim , Thagata ; Java , Tcibodas; Pengalengan, 1300 m).

— Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Cameron, 1928d: 92 ( Eleusis ; illustrations; Sumatra).

— Bernhauer, 1928: 1 ( Eleusis ; Sumatra).

— Cameron, 1930: 82 ( Eleusis ; characters; Burma; Assam; Java; Lombok).

— Scheerpeltz, 1933: 999 ( Eleusis ; catalog).

— Abdullah and Qadri, 1970: 118 ( Eleusis ; characters; East Pakistan).

- DISTRIBUTION: India, Bangladesh, Myanmar, Indonesia.

nova Bernhauer, 1926: 19 ( Eleusis ; Type locality: Nordost-Indien: Kobo, 400 ft.).

— Cameron, 1930: 82 ( Eleusis ; synonym of viridans ).

— Scheerpeltz, 1933: 998 ( Eleusis ; synonym of viridans ).

warburgi Fauvel, 1904b: 82 ( Eleusis ; Type locality: Celebes: Minahassa ).

— Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

- DISTRIBUTION: Indonesia.

Renardia

[5 species; Nearctic and Neotropical Regions]

Renardia Motschulsky, 1865: 583 (species included: jubilaea ). TYPE SPECIES: Renardia jubilaea Motschulsky , fixed by monotypy.

— Leng, 1920: 95 (catalog of North American species).

— Scheerpeltz, 1933: 1072 (world catalog supplement).

— Blackwelder, 1952: 338 (type species: jubilaea ).

— Moore and Legner, 1974b: 82 (characters; notes).

— Moore and Legner, 1975: 170 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 148 (characters; notes).

— Downie and Arnett, 1996: 428 (characters; key to species of Northeastern North America).

— Newton, Thayer, Ashe, and Chandler, 2000: 377 (2 Nearctic species; notes; characters in key). Eumalus Sharp, 1887: 732 (species included: strigosus ; nigriceps ). TYPE SPECIES: Eumalus nigriceps Sharp , fixed by subsequent designation by Lucas, 1920: 281.

— Bernhauer and Schubert, 1910: 10 (world catalog; 3 species).

— Leng, 1920: 93 (catalog of North American species).

— Scheerpeltz, 1933: 996 (world catalog supplement).

— Blackwelder, 1942: 88 (type species: strigosus ).

— Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies).

— Steel, 1950a: 215 (characters in key).

— Blackwelder, 1952: 156 (type species).

— Hatch, 1957: 244 (characters; species of Pacific Northwest).

— Hammond, 1970: 69, 70 (synonym of Renardia ).

canadensis Horn, 1871: 297 , 299 ( Isomalus ; Type locality: Grimsby , Ontario).

— Bernhauer and Schubert, 1910: 10 ( Eumalus ; synonym of nigrella ).

— Moore and Legner, 1975: 170 ( Renardia ; cited as valid species; catalog).

— Downie and Arnett, 1996: 428 ( Renardia ; characters; USA).

- DISTRIBUTION: Canada.

nigrella LeConte, 1863a: 59 ( Isomalus ; Type locality: California, San Jose and Fort Tejon ).

— Horn, 1871: 297, 298, 299 ( Isomalus ; characters; California).

— Fauvel, 1878d: 182 [= 1878e: 18] ( Eleusis ; characters; Canada; California; Texas).

— Sharp, 1887: 733 ( Eumalus ).

— Bernhauer and Schubert, 1910: 10 ( Eumalus ; catalog).

— Hatch, 1957: 244 ( Eumalus ; characters; British Columbia; Washington; Oregon; Idaho).

— Legner and Moore, 1977: 174 ( Renardia ; found under bark).

— Downie and Arnett, 1996: 428 ( Renardia ; characters; Canada; USA).

- DISTRIBUTION: Canada, USA.

jubilaea Motschulsky, 1865: 584 ( Renardia ; Type locality: environs de New York; [Note: See lectotype designation by Hammond, 1970]).

— Scheerpeltz, 1933: 1072 ( Renardia ; catalog).

— Hammond, 1970: 69, 70 ( Renardia ; lectotype designation; notes; synonym of nigrella ). jubilaea Motschulsky, 1865 , see: nigrella LeConte, 1863 .

nigriceps Sharp, 1887: 733 ( Eumalus ; Type locality: Guatemala, Capetillo ).

— Bernhauer and Schubert, 1910: 10 ( Eumalus ; catalog).

- DISTRIBUTION: Guatemala.

strigosus Sharp, 1887: 733 ( Eumalus ; Type locality: Guatemala, Totonicapam 8500 to

10,500 feet).

— Bernhauer and Schubert, 1910: 10 ( Eumalus ; catalog).

tenenbaumi Bernhauer, 1929 h: 186 ( Eumalus ; Type locality: Mexiko: Desierto de los Leones). — Scheerpeltz, 1933: 996 ( Eumalus ; catalog).

- DISTRIBUTION: Mexico.

Zeoleusis

[12 species; Neotropical, Oriental, and Australian Regions]

Zeoleusis Steel, 1950a: 215 (species included: virgula , semirufus , insularis , sororcula , polita , annae , parva , banghaasi , helferi , exigua , fusciceps ). TYPE SPECIES: Eleusis virgula Fauvel , fixed by original designation.

— Steel, 1950a: 219 (key to species).

— Coiffait and Saiz, 1968: 462 (characters; key to Chilean species).

— Newton, 1985: 203 (notes on austral distribution).

— Newton, 1990: 91 (characters in key; two described Australian species).

annae Steel, 1950a: 225 ( Zeoleusis ; Type locality: New South Wales: Sydney ).

— Newton, 1990: 91 ( Zeoleusis ).

- DISTRIBUTION: Australia.

banghaasi Bernhauer, 1926d: 253 ( Eleusis ; Type locality: Siargao, Dapa. Palawan, Binaluan. Luzon, Provinz Mountain, Balbalasang. Mindanao, Provinz Lanao , Mumungan. Masbate, Aroroy ).

— Scheerpeltz, 1933: 996 ( Eleusis ; catalog).

— Steel, 1950a: 231 ( Zeoleusis ; characters; Philippines Islands).

- DISTRIBUTION: Philippines.

decipiens Cameron, 1932 , see: exigua Bernhauer, 1926 .

exigua Bernhauer, 1926d: 253 ( Eleusis ; variety of banghaasi ; Type locality: Not cited).

— Scheerpeltz, 1933: 996 ( Eleusis ; synonym of augustae ).

— Steel, 1950a: 233 ( Zeoleusis ; cited as valid species; characters; Philippines; Malaysia).

- DISTRIBUTION: Philippines, Malaysia.

decipiens Cameron, 1932a: 124 ( Eleusis ; Type locality: Pahang: Fraser’s Hill, alt. 4000 feet). — Steel, 1950a: 232 ( Zeoleusis ; synonym of exigua ).

fusciceps Kraatz, 1859: 184 ( Isomalus ; Type locality: Ceylan).

— Bernhauer and Schubert, 1910: 11 ( Eleusis ; catalog).

— Cameron, 1921a: 362, 398 ( Eleusis ; characters; Singapore).

— Bernhauer, 1926d: 253 ( Eleusis ; characters; Philippines).

— Cameron, 1930: 88 ( Eleusis ; characters; Ceylon; Andamans; Siwaliks; Singapore; Sumatra).

— Cameron, 1930b: 160 ( Eleusis ; British North Borneo).

— Scheerpeltz, 1933: 997 ( Eleusis ; catalog).

— Steel, 1950a: 233 ( Zeoleusis ; characters; Ceylon; India; Singapore; Sumatra; Philippines).

— Jarrige, 1963a: 120 ( Eleusis ; Ceylon).

— Abdullah and Qadri, 1970: 116 ( Eleusis ; characters; West Pakistan).

— Coiffait, 1981f: 331 ( Eleusis ; cited as fuscipes ; Andaman Islands).

— Naomi, 1996: 51 ( Zeoleusis ; Japan).

- DISTRIBUTION: Pakistan, India, Bangladesh, Sri Lanka , Andaman Islands, Borneo, Singapore, Indonesia, Philippines, Japan.

helferi Steel, 1950a: 230 ( Zeoleusis ; Type locality: Bosnek , Biak Island, New Guinea).

- DISTRIBUTION: New Guinea.

insularis Cameron, 1936: 3 ( Eleusis ; Type locality: Rapa, Mount Vaitau, altitude 1100 feet). — Steel, 1950a: 221 ( Zeoleusis ; characters; Rapa Island).

- DISTRIBUTION: Austral Islands.

myrmidon Fairmaire and Germain, 1861: 455 ( Isomalus ; Type locality: Santiago).

— Fauvel, 1867a: 53 [= 1868: 58] ( Isomalus ; characters; Chile).

— Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Coiffait and Saiz, 1968: 463 ( Zeoleusis ; characters; Chile).

parva Blackburn, 1892: 24 ( Eleusis ; Type locality: New South Wales, Blue Mountains ).

— Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Steel, 1950a: 224 ( Zeoleusis ; characters; Australia).

- DISTRIBUTION: Australia.

polita Steel, 1950a: 227 ( Zeoleusis ; Type locality: Ceylon).

- DISTRIBUTION: Sri Lanka.

semirufa Fairmaire and Germain, 1861: 455 ( Isomalus ; Type locality: Ile de Juan Fernandez). — Fauvel, 1867a: 53 [= 1868: 58] ( Isomalus ; characters; Chile).

— Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Bernhauer, 1921a: 41 ( Eleusis ; Juan Fernandez).

— Scheerpeltz, 1933: 998 ( Eleusis ; catalog).

— Steel, 1950a: 223 ( Zeoleusis ; characters; Juan Fernandez).

— Coiffait and Saiz, 1968: 463 ( Zeoleusis ; characters; Chile).

- DISTRIBUTION: Chile, Juan Fernandez.

sororcula Cameron, 1936: 4 ( Eleusis ; Type locality: Rapa, Mount Vaitau, altitude, 1100 feet). — Steel, 1950a: 222 ( Zeoleusis ; characters; Rapa Island).

- DISTRIBUTION: Austral Islands.

virgula Fauvel, 1889b: 243 ( Eleusis ; Type locality: Nouvelle-Zelande: Auckland; Greymouth). — Bernhauer and Schubert, 1910: 12 ( Eleusis ; catalog).

— Steel, 1950a: 228 ( Zeoleusis ; characters; New Zealand).

— Kuschel, 1990: 26 ( Zeoleusis ; endemic; winged; habitat; New Zealand).

— Emberson, 1998: 35 ( Zeoleusis ; collecting notes; Chatham Islands).

- DISTRIBUTION: New Zealand, Chatham Islands.

Tribe Leptochirini

Leptochirini Sharp, 1887 .

— Eichelbaum, 1909: 110 (list of genera with some references; generic synonyms; number of genera; number of species in each genus; general distribution of genera).

— Cameron, 1930: 89 (characters; key to British Indian genera).

— Blackwelder, 1942: 79 (characters).

— Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies).

— Newton and Thayer, 1992: 59 (tribe of Osoriinae ).

Leptochiri Sharp, 1887.

— Bernhauer and Schubert, 1910: 13 (world catalog; Leptochirus , Borolinus , Priochirus, Thoracochirus ).

— Scheerpeltz, 1933: 999 (world catalog supplement).

Leptochirina Sharp, 1887: 733 (genera included: Leptochirus , Priochirus ). TYPE GENUS: Leptochirus (see Newton and Thayer, 1992: 59).

— Jakobson, 1908: 446, 466 (characters; list of species of Russia and western Europe).

Borolinus

[12 species; Palaearctic, Oriental, Australian, and Oceanic Regions] Borolinus Bernhauer, 1903: 116 , 133 (species included: semirufus , antilope , minutus , javanicus , borneensis ). TYPE SPECIES: Leptochirus javanicus Laporte , fixed by subsequent designation by R. Lucas, 1920: 144.

— Bernhauer and Schubert, 1910: 15 (world catalog; 5 species).

— Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog).

— Cameron, 1925: 4 (catalog of Indian species).

— Cameron, 1930: 92 (characters; key to British Indian species).

— Scheerpeltz, 1933: 1001 (world catalog supplement).

— Blackwelder, 1952: 84 (type species). — Shibata, 1973c: 25 (checklist of species of Taiwan).

— Shibata, 1976: 87 (checklist of species of Japan).

— Hammond, 1984: 196 (checklist of species of Borneo).

— Drugmand, 1998a: 206 (list of types in l’Institut royal des Sciences naturelles de Belgique). antilope Fauvel, 1895: 181 ( Leptochirus ; Type locality: Bírmanie, Carin Ghécu, 1300– 1400 m; Senmijion).

— Heller, 1898: 6 ( Priochirus ; checklist).

— Bernhauer, 1903: 134 ( Borolinus ; characters).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; catalog).

— Cameron, 1928d: 94 ( Borolinus ; Sumatra).

— Cameron, 1930: 93 ( Borolinus ; characters; Burma).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog; Siam).

- DISTRIBUTION: Myanmar, Thailand, Indonesia.

nigripennis Fauvel, 1902: 10 ( Leptochirus ; subgenus Priochirus ; variety of antilope ; Type locality: Sumatra: Si-Rambé).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; synonym of antilope ).

— Cameron, 1928d: 94 ( Borolinus ; variety of antilope ; Sumatra).

— Cameron, 1936a: 26 ( Leptochirus ; subgenus Strongylochirus ; variety of antilope ; Java). bispinus Erichson, 1840 , see: minutus Laporte, 1840 .

borneensis Bernhauer, 1903: 133 ( Borolinus ; Type locality: Borneo).

— Fauvel, 1905e: 77 ( Leptochirus ; synonym of minutus ).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; catalog).

— Wendeler, 1928c: 117 ( Borolinus ; Philippines).

- DISTRIBUTION: Borneo, Philippines.

cruentus Fauvel, 1895 , see: minutus Laporte, 1840 .

curticollis Bernhauer, 1926b: 38 ( Borolinus ; Type locality: Philippinen: Mindanao, Iligan). — Wendeler, 1928c: 117 ( Borolinus ; Philippines).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

- DISTRIBUTION: Philippines.

furcatus Wendeler, 1928c: 119 ( Borolinus ; Type locality: Philippinen: Luzon : Imugan,

Mt. Isarog).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

- DISTRIBUTION: Philippines.

inermis Fauvel, 1902 , see: minutus Laporte, 1840 .

javanicus Laporte, 1835: 126 ( Leptochirus ; Type locality: Java).

— Erichson, 1840: 827 ( Leptochirus ; characters; Java).

— Fauvel, 1886b: 144 ( Leptochirus ; Philippines; Java; Sumatra).

— Heller, 1898: 7 ( Priochirus ; checklist).

— Bernhauer, 1903: 134 ( Borolinus ; characters).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; catalog).

— Cameron, 1928d: 94 ( Borolinus ; Sumatra).

— Bernhauer, 1928: 1 ( Borolinus ; Sumatra).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

— Frank, 1982: 6 ( Borolinus ; list of parasites).

- DISTRIBUTION: Philippines, Indonesia.

nigricollis Fauvel, 1902: 10 ( Leptochirus ; subgenus Priochirus ; variety of javanicus ; Type locality: nord de Luçon).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; synonym of javanicus ).

— Scheerpeltz, 1933: 1001 ( Borolinus ; synonym of javanicus ).

minutus Laporte, 1840: 186 ( Leptochirus ; Type locality: Java).

— Fauvel, 1878f: 186 ( Leptochirus ; characters; New Guinea; Java; Borneo; Philippines).

— Fauvel, 1886b: 144 ( Leptochirus ; Philippines; Siam; Malacca; Borneo; Java; Gilolo; New Guinea).

— Heller, 1898: 8 ( Priochirus ; checklist).

— Bernhauer, 1903: 134 ( Borolinus ; characters).

— Fauvel, 1905e: 77 ( Leptochirus ; Java).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; catalog).

— Bernhauer, 1922c: 221 ( Borolinus ; Formosa).

— Cameron, 1928d: 94 ( Borolinus ; Sumatra).

— Bernhauer, 1928: 1 ( Borolinus ; Sumatra).

— Wendeler, 1928c: 117 ( Borolinus ; Philippines).

— Cameron, 1930: 94 ( Borolinus ; characters; Assam; Sikkim; Burma; Siam; Sumatra; Java; Lombok; Borneo).

— Cameron, 1930b: 160 ( Priochirus ; British North Borneo).

— Cameron, 1931: 355 ( Borolinus ; New Britain).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

— Cameron, 1936a: 26 ( Leptochirus ; subgenus Strongylochirus ; Java).

— Cameron, 1937a: 84 ( Borolinus ; New Guinea).

— Paulian, 1941: 132 ( Borolinus ; larval characters; India).

— Cameron, 1949b: 456 ( Borolinus ; China).

— Scheerpeltz, 1965: 98 ( Borolinus ; Burma).

— Abdullah and Qadri, 1970: 116 ( Borolinus ; characters; East Pakistan).

— Coiffait, 1981f: 332 ( Borolinus ; Andaman Islands).

— Frank, 1982: 6 ( Borolinus ; list of parasites).

— Biswas and Biswas, 1985: 142 ( Borolinus ; India).

- DISTRIBUTION: China, Taiwan, New Guinea, New Britain, Philippines, Borneo, Indonesia, Singapore, Malaysia, Thailand, Myanmar, Andaman Islands, Bangladesh, India.

bispinus Erichson, 1840: 827 ( Leptochirus ; Type locality: Java).

— Fauvel, 1878f: 186 ( Leptochirus ; synonym of minutus ).

— Fauvel, 1905e: 77 ( Leptochirus ; synonym of minutus ).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; synonym of minutus ).

— Cameron, 1930: 94 ( Borolinus ; synonym of minutus ).

— Scheerpeltz, 1965: 98 ( Borolinus ; synonym of minutus ).

cruentus Fauvel, 1895: 181 ( Leptochirus ; variety of minutus ; Type locality: Malacca; Singapore; Sumatra; Borneo).

— Heller, 1898: 8 ( Priochirus ; variety of minutus ; checklist).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; synonym of minutus ).

— Cameron, 1921a: 362, 399 ( Borolinus ; variety of minutus ; characters; Singapore).

— Cameron, 1928d: 94 ( Borolinus ; variety of minutus ; Sumatra).

— Scheerpeltz, 1933: 1001 ( Borolinus ; variety of minutus ).

inermis Fauvel, 1902: 10 ( Leptochirus ; subgenus Priochirus ; variety of minutus ; Type locality: Sumatra: Palembang).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; synonym of minutus ).

nigricollis Fauvel, 1902 , see: javanicus Castelnau, 1835 .

nigripennis Fauvel, 1902 , see: antilope Fauvel, 1895 .

oceani Bernhauer, 1919: 360 ( Borolinus ; Type locality: Philippinen: Magdalena , Laguna).

— Wendeler, 1928c: 117 ( Borolinus ; Philippines).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

- DISTRIBUTION: Philippines.

rufus Cameron, 1928 g: 424 ( Borolinus ; Type locality: Borneo: Mount Dulit, 3500 feet).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

- DISTRIBUTION: Borneo.

samarensis Bernhauer, 1928: 2 ( Borolinus ; Type locality: Island Samar).

semirufus Fauvel, 1895: 181 ( Leptochirus ; Type locality: Bírmanie, Carin Cheba , 900– 1100 m; Asciuii Cheba).

— Heller, 1898: 8 ( Priochirus ; checklist).

— Bernhauer, 1903: 134 ( Borolinus ; characters).

— Bernhauer and Schubert, 1910: 15 ( Borolinus ; catalog).

— Cameron, 1930: 93 ( Borolinus ; characters; Burma).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

- DISTRIBUTION: Myanmar.

sikkimensis Bernhauer, 1919: 359 ( Borolinus ; Type locality: Himalaya : Sikkim).

— Cameron, 1930: 94 ( Borolinus ; characters; Sikkim; Burma).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

— Scheerpeltz, 1965: 98 ( Borolinus ; Burma).

— Abdullah and Qadri, 1970: 116 ( Borolinus ; characters; East Pakistan).

— Biswas and Biswas, 1985: 142 ( Borolinus ; India).

- DISTRIBUTION: India, Bangladesh, Myanmar.

sumatrensis Bernhauer, 1928: 2 ( Borolinus ; Type locality: Sumatra: Brastagi ).

— Scheerpeltz, 1933: 1001 ( Borolinus ; catalog).

- DISTRIBUTION: Indonesia.

Leptochirus

[61 species; Neotropical, Palaearctic, and Oriental Regions]

Leptochirus Germar, 1824: 35 (species included: scoriaceus ). TYPE SPECIES: Leptochirus scoriaceus Germar , fixed by monotypy.

— Latreille, 1825: 245 (cited as Leptochire; synonym of Zirophore).

— Berthold, 1827: 332 (characters).

— Erichson, 1840: 824 (characters).

— Laporte, 1840: 186 (characters).

— Lacordaire, 1854: 125 (characters; notes; list of species).

— Fauvel, 1864b: 8 (characters; notes; list of species; larval characters).

— Fauvel, 1865: 12 (characters; notes; list of species; larval characters).

— Gemminger and Harold, 1868: 678 (catalog).

— Sharp, 1876: 420 (characters; notes).

— Fauvel, 1877: 184 (notes; key to species).

— Duvivier, 1883: 205 (catalog).

— Sharp, 1887: 733 (characters; Mexican and Central species).

— Heller, 1898: 5, 6, 8 (discussion; checklist of and key to species).

— Bernhauer, 1903: 116, 117 (characters; key to species).

— Bernhauer and Schubert, 1910: 13 (world catalog; 35 species).

— Sharp and Muir, 1912: 501 (characters of male genitalia).

— Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog).

— Cameron, 1925: 3 (catalog of Indian species).

— Cameron, 1930: 89 (characters; key to British Indian species).

— Scheerpeltz, 1933: 999 (world catalog supplement).

— Paulian, 1941: 128 (larval characters).

— Blackwelder, 1943: 162 (characters; type species: scoriaceus ; key to species of West Indies).

— Blackwelder, 1944: 109 (checklist of species from Mexico, Central and South America, and the West Indies).

— Blackwelder, 1952: 215 (type species: scoriaceus ).

— Hammond, 1984: 196 (checklist of species of Borneo).

— Drugmand, 1998a: 212 (list of types in l’Institut royal des Sciences naturelles de Belgique).

Mesochirus Bernhauer, 1903: 120 (subgenus of Leptochirus ; species included: extimus , maxillosus , latro , brunneoniger , cayennensis , arcifer , tenuis ). TYPE SPECIES: Cucujus maxillosus Fabricius , fixed by subsequent designation by R. Lucas, 1920: 405.

— Bernhauer and Schubert, 1910: 14 (catalog; subgenus of Leptochirus ).

Tropiochirus Bernhauer, 1903: 120 (subgenus of Leptochirus ; species included: helleri , proteus ). TYPE SPECIES: Leptochirus proteus Fauvel , fixed by subsequent designation by R. Lucas, 1920: 659.

— Bernhauer and Schubert, 1910: 14 (catalog; subgenus of Leptochirus ).

— Scheerpeltz, 1933: 1000 (world catalog supplement; subgenus of Leptochirus ).

— Blackwelder, 1943: 162 (subgenus of Leptochirus ; type species).

— Blackwelder, 1952: 399 (subgenus of Leptochirus ; type species).

Strongylochirus Bernhauer, 1903: 120 (subgenus of Leptochirus ; species included: quadridens , loriae , laevis , atkinsoni ). TYPE SPECIES: Leptochirus laevis Laporte , fixed by subsequent designation by Lucas, 1920: 618.

— Bernhauer and Schubert, 1910: 14 (catalog; subgenus of Leptochirus ).

— Scheerpeltz, 1933: 1000 (world catalog supplement; subgenus of Leptochirus ).

— Blackwelder, 1943: 162 (subgenus of Leptochirus ; type species laevis ).

— Blackwelder, 1952: 364 (type species).

— Biswas and Sengupta, 1977: 341 (subgenus of Leptochirus ; key to species of India).

albertisi Fauvel, 1878 , see: Priochirus .

alternus Fauvel, 1878 , see: Priochirus .

alticola Bernhauer, 1912d: 28 ( Leptochirus ; Type locality: Columbia: Cali: S. Antonio, 2000 m; Rio Aquatal, 1880 m).

— Scheerpeltz, 1933: 999 ( Leptochirus ; catalog).

- DISTRIBUTION: Colombia.

andinus Bernhauer, 1917f: 46 ( Leptochirus ; subgenus Leptochirus ; Type locality: WestKolumbien: Pacho in den Ost-Kordilleren, 1900 m).

— Scheerpeltz, 1933: 999 ( Leptochirus ; catalog).

- DISTRIBUTION: Colombia.

angustulus Sharp, 1887: 736 ( Leptochirus ; Type locality: Colombia, Venezuela).

— Heller, 1898: 6 ( Leptochirus ; checklist).

— Bernhauer, 1903: 121 ( Leptochirus ; characters).

— Bernhauer, 1903: 122 ( Leptochirus ; characters).

— Bernhauer and Schubert, 1910: 13 ( Leptochirus ; subgenus Leptochirus ; catalog).

— Scheerpeltz, 1933: 999 ( Leptochirus ; subgenus Leptochirus ; catalog).

- DISTRIBUTION: Colombia, Venezuela.

antennarius Fauvel, 1878 , see: Priochirus .

anthracinus Fauvel, 1902 , see: maxillosus Fabricius, 1801 .

antilope Fauvel, 1895 , see: Borolinus .

apicalis Eppelsheim, 1895 , see: Priochirus .

arcifer Fauvel, 1902: 17 ( Leptochirus ; Type locality: Bolivie: Yuracaris , Yungas; Pérou: Marcapata).

— Bernhauer, 1903: 125, 131 ( Leptochirus ; characters; Peru; Bolivia).

— Bernhauer and Schubert, 1910: 14 ( Leptochirus ; subgenus Mesochirus ; catalog).

- DISTRIBUTION: Bolivia, Peru.

ascendens Fauvel, 1903 , see: Priochirus .

aspericollis Fauvel, 1902 , see: Thoracochirus .

atkinsoni Fauvel, 1895: 182 ( Leptochirus ; Type locality: Bírmanie, Carin Cheba , 900–1400 m; Tenasserim, Tavoy).

— Heller, 1898: 6 ( Leptochirus ; checklist).

— Bernhauer, 1903: 126 ( Leptochirus ; characters).

— Bernhauer and Schubert, 1910: 14 ( Leptochirus ; subgenus Strongylochirus ; catalog).

— Cameron, 1930: 92 ( Leptochirus ; subgenus Strongylochirus ; characters; Assam; Burma; Siam). — Scheerpeltz, 1933: 1000 ( Leptochirus ; subgenus Strongylochirus ; catalog).

— Biswas and Sengupta, 1977: 345 ( Leptochirus ; subgenus Strongylochirus ; characters; Burma; India).

— Biswas and Biswas, 1985: 142 ( Leptochirus ; subgenus Strongylochirus ; India).

- DISTRIBUTION: India, Bangladesh, Myanmar, Thailand.

beccarii Fauvel, 1878 , see: Priochirus .

bernhaueri Wendeler, 1957a: 47 ( Leptochirus ; replacement name for sharpi Bernhauer ).

- DISTRIBUTION: Mexico.

mexicanus Sharp, 1887: 739 ( Leptochirus ; [preoccupied]; Type locality: Mexico, Cordova). — Bernhauer and Schubert, 1910: 14 ( Leptochirus ; synonym of sharpi ).

— Wendeler, 1957a: 47 ( Leptochirus ; synonym of bernhaueri ).

sharpi Bernhauer, 1903: 123 , 130 ( Leptochirus ; [preoccupied; replacement name for mexicanus Sharp ; Note: Erroneously cited by Bernhauer and Schubert, 1910: 14

as a subsequent reference of sharpi Fauvel ]).

— Wendeler, 1957a: 47 ( Leptochirus ; synonym of bernhaueri ).

bicolor P. Lucas, 1857 , see: maxillosus Fabricius, 1801 .

bicornis Fauvel, 1864 , see: Priochirus .

bifoveatus Eppelsheim, 1895 , see: Priochirus .

bifurcatus Fauvel, 1879 , see: Priochirus .

bipunctatus Fauvel, 1895 , see: Priochirus .

bispinus Erichson, 1840 , see: Borolinus .

bolivianus Heller, 1898 , see: maxillosus Fabricius, 1801 .

brachycerus Kraatz, 1859 , see: Priochirus .

brasilianus Bernhauer, 1903: 121 , 128 ( Leptochirus ; Type locality: Brasilia).

— Bernhauer and Schubert, 1910: 13 ( Leptochirus ; subgenus Leptochirus ; catalog).