Zadadra confusa Dubatolov, Volynkin, N. Singh, Joshi & Černý,

Dubatolov, Vladimir V., Volynkin, Anton V., Singh, Navneet, Joshi, Rahul & Černý, Karel, 2021, On the taxonomy of the Prabhasa / Zadadra generic complex with descriptions of two new genera and two new species (Lepidoptera, Erebidae, Arctiinae, Lithosiini), Zootaxa 4966 (5), pp. 519-534: 529-534

publication ID

https://doi.org/10.11646/zootaxa.4966.5.2

publication LSID

lsid:zoobank.org:pub:A23BF4A7-252F-4099-BAE8-E3EFC00BDCD8

DOI

http://doi.org/10.5281/zenodo.4745269

persistent identifier

http://treatment.plazi.org/id/03E787A2-FFE9-B250-FF73-FCBBFA68FEA9

treatment provided by

Plazi

scientific name

Zadadra confusa Dubatolov, Volynkin, N. Singh, Joshi & Černý
status

sp. nov.

Zadadra confusa Dubatolov, Volynkin, N. Singh, Joshi & Černý  , sp. nov.

( Figs 15–18View FIGURES 11–18, 29, 30View FIGURES 27–30, 32View FIGURES 31–32, 38View FIGURES 33–38)

Type material. Holotype ( Figs 15View FIGURES 11–18, 29View FIGURES 27–30): ♂, “ Nepal, Ganesh Himal , 2 km N of Dhunche, 2050m, 15.IX.1995, leg. Lajos Németh ”, gen. slide No.: ZSM Arct. 2019-508 (prepared by Volynkin) (MWM/ ZSM). 

Paratypes. NEPAL: 5 ♂, 1 ♀, the same data as in holotype (MWM/ ZSM)  ; 1 ♂, 3 ♀, Ganesh Himal , 2 km W of Thangjet, 85°17’E, 28°10’N, 2300m, 23.IX.1994, leg. Márton Hreblay & Tibor Csóvári (MWM/ ZSM)GoogleMaps  ; 11 ♂, 3 ♀, Ganesh Himal , 2 km E of Thangjet, 85’19’E, 28’10’N, 2165m, 17.IX.1994, leg. Márton Hreblay & Tibor Csővári (MWM/ ZSM)  ; 2 ♂, 1 ♀, 2165m, Ganesh Himal , 2 km E of Thangjet, 16.X.1995, 85’19’E, 28’11’N, leg. Hreblay & Bodi (MWM/ ZSM)  ; 2 ♂, 6 ♀, Ganesh Himal , 2 km E of Thangjet, 2165m, 16–17.IX.1995, leg. Lajos Németh (MWM/ ZSM)  ; 14 ♂, 61 ♀, Ganesh Himal , 1700m, 3 km W Gogne, 85°12’E, 28°05.5’N, 23.IX.1995, leg. B. Herczig & Gy. M. László (MWM/ ZSM)GoogleMaps  ; 22 ♂, 28 ♀, Ganesh Himal , 2200m, 4 km SW Haku, 85°15.5’E, 28°06.5’N, 22.IX.1995, leg. B. Herczig & Gy. M. László (MWM/ ZSM)GoogleMaps  ; 2 ♀, Ganesh Himal, 1040m, Mailung Khola , 85°12.5’E, 28°04.5N, 24.X.1995, leg. L. Peregovits & L. Ronkay (MWM/ ZSM)GoogleMaps  ; 3 ♂, 2 ♀, Ganesh Himal, 2520m, near Godlang , 85°17’E, 28°10’N, 13.X.1995, leg. L. Peregovits & L. Ronkay (MWM/ ZSM)GoogleMaps  ; 1 ♂, 1 ♀, Ganesh Himal , 2420m, 2 km W Gholjong, 85°18’E, 28°11’N, 12.X.1995, leg. S. Kovács, gen. slide No.: ZSMGoogleMaps  Arct. 2019-509 (male) (prepared by Volynkin) (MWM/ ZSM)  ; 5 ♂, 8 ♀, Ganesh Himal, 2720m, above Nesim , 85°16’E, 28°08.5’N, 21.IX.1995, leg. B. Herczig & Gy. M. László, gen. slide No.: ZSMGoogleMaps  Arct. 2019-510 (male) (prepared by Volynkin) (MWM/ ZSM)  ; 4 ♂, 3 ♀, Ganesh Himal , 3 km NE of Sunpati, 2330m, 13. V  .1993, leg. M. Hreblay & G. Csorba (MWM/ ZSM); 2 ♂, Ganesh Himal, 1040m, Mailung Khola , 85°12.5’E, 28°04.5N, 24.X.1995, leg. L. Peregovits & L. Ronkay (MWM/ ZSM)GoogleMaps  ; 3 ♂, 4 ♀, Ganesh Himal, valley of Trisuli River , 2 km S of Betrawati, 930m, 25.IX.1995, leg. L. Németh (MWM/ ZSM)  ; 1 ♂, Ganesh Himal, Syabrubesi , 1520m, 12. VI  .1993, leg. M. Hreblay, G. Csorba (MWM/ ZSM); 2 ♀, Ganesh Himal, 2 km W of Gadrang, Oak-forest , 2720m, 18–19.IX.1995, leg. L. Németh (MWM/ ZSM)  ; 1 ♂, Koshi, Terhathum area, Chitre , 2500m, 87°24’E 27°05’N, 17.X.1996, leg. Gy. M. László and G. Ronkay, gen. slide No.: ZSM Arct. 2019-511 (prepared by Volynkin) (MWM/ ZSM)GoogleMaps  ; 61 ♂, 38 ♀, Langtang, 1950m, 1.5 km NE Dhunche , 85°18’E, 28°06’N, 24.IX.1994, leg. Csorba & Ronkay (MWM/ ZSM)GoogleMaps  ; 9 ♂, Langtang 5 km NNE of Dunche, Barkhu , 85°18’E, 28°08’N, 1835m, 16.IX.1994, leg. Márton Hreblay & Tibor Csóvári (MWM/ ZSM)GoogleMaps  ; 3 ♂, 8 ♀, Langtang , 2110m, 9 km S Dhunche, 85°14’E, 28°04’N, 23.IX.1994, leg. Csorba & Ronkay (MWM/ ZSM)GoogleMaps  ; 4 ♂, 9 ♀, 1835m, Langtang 5 km NNE of Dunche, Barkhu , 85’18’E, 28’08’N, leg. M. Hreblay & T  . Csővári , 16.IX.1994 (MWM/ ZSM)  ; 1 ♂, 1 ♀, Mechi, Taplejung Area, Nesum village , 1550m, 87°29’E, 27°17’N, 21.X.1996, leg. Gy. M. László & G. Ronkay, gen. slide Nos.: MWM 31606View Materials (male) and ZSMGoogleMaps  Arct. 2019-516 (female) (prepared by Volynkin) (MWM/ ZSM)  ; 5 ♂, 9 ♀, Annapurna Himal, Ulleri , 1900m, 83°43’E, 28°23’N, 3.X.1994, leg. Csorba, Ronkay (MWM/ ZSM)GoogleMaps  ; 1 ♀, Annapurna Himal, valley of Kali Gandaki, 1300m, near Tatopani , 83°39’E, 28°29’N, 03. VIGoogleMaps  .1996, leg. Gy. M. László and G. Ronkay (MWM/ ZSM); 57 ♂, 116 ♀, Annapurna Himal, valley of Kali Gandaki, Kokethanti village , 2650m, 17. VI.1996, leg. Gy. M. László and G. Ronkay, gen. slide Nos.: ZSM Arct. 2019-507 (male) and 2019-515 (female) (prepared by Volynkin) (MWM/ ZSM)  ; 1 ♂, 3 ♀, Annapurna Himal, valley of Kali Gandaki, 3 km NE Tukuche, 2750m, 16. VI  .1996, leg. Gy. M. László and G. Ronkay (MWM/ ZSM); 31 ♂, 34 ♀, Annapurna Himal, valley of Kali Gandaki, 2080m, near Ghasa , 83°39.5’E, 28°36’N, 04. VI.1996, leg. Gy. M. László and G. Ronkay, gen. slide No.: ZSM Arct. 2019-506 (male) (prepared by Volynkin) (MWM/ ZSM)GoogleMaps  ; 43 ♂, 91 ♀, the same locality and collectors, 18–19. VI  .1996, gen. slide No.: ZSM  Arct. 2019-514 (female) (prepared by Volynkin) (MWM/ ZSM)  ; 10 ♂, 19 ♀, Annapurna Himal , 1700m, 1 km N of Tal, 84’23’’E, 28’28’’N, 08. VI  .1996, leg. Hreblay & Szaboky (MWM/ ZSM); 1 ♂, 2 ♀, Annapurna Himal, Banthanti village , 2420m, 83°43’E, 28°22.5’N, 01. VIGoogleMaps  .1996, leg. Gy. M. László and G. Ronkay (MWM/ ZSM); 1 ♂, Annapurna Himal, Thini , 1 km S Jomsom, 3000m, 83°44’E, 28°46’N, 06–07. VIGoogleMaps  .1996, leg. Gy. M. László and G. Ronkay (MWM/ ZSM); 1 ♂, the same locality and collectors, 15. VIGoogleMaps  .1996 (MWM/ ZSM)GoogleMaps  ; 3 ♀, Annapurna Himal, 1200m, Thini , 1 km N of Syange, 84°25’E, 28°24’N, 07. VIGoogleMaps  .1996, leg. Hreblay & Szaboky (MWM/ ZSM); 1 ♂, 10 ♀, Annapurna Himal, 1950m, Talbagar , 83°39’E, 28°34’N, 24. VIGoogleMaps  .1996, leg. Hreblay & Szaboky (MWM/ ZSM); 1 ♀, Annapurna Himal, Nangethanti , 2500m, 83°43’E, 28°23’N, 4.X.1994, leg. Csorba & Ronkay (MWM/ ZSM)GoogleMaps  ; 3 ♀, Annapurna Himal, Bhaleodar , 2400m, 2 km SE Nangethanti, 83°44’E, 28°23’N, 8.X.1994, leg. Csorba & Ronkay (MWM/ ZSM)GoogleMaps  ; 1 ♀, Annapurna Himal , 8 km SW of Tatopani, 1200m, 31.VIII.1996, 83°37’E, 28°27’N, leg. Chenga Sherpa (MWM/ ZSM)GoogleMaps  ; 2 ♀, Annapurna Himal , 850m, 1 km N of Besisahar, 84°23’E, 28°14’N, 05. VIGoogleMaps  .1996, leg. Hreblay & Szaboky (MWM/ ZSM); 1 ♀, Annapurna Himal, 2200m, Bagarchhap , 84°20’E, 28°32’N, 09. VIGoogleMaps  .1996, leg. Hreblay & Szaboky (MWM/ ZSM); 3 ♂, 13 ♀, valley of Tamea Kosi River , 5 km S of Piguti, 950m, 8–9.X.1995, leg. L. Németh (MWM/ ZSM)  ; 2 ♂, 8 ♀, valley of Tamea Kosi River , 1 km N of Dolakha, 1700m, 12.X.1995, leg. L. Németh (MWM/ ZSM)  ; 1 ♀, valley of Tamea Kosi River , 3 km N of Malephu, 900m, 10–11.X.1995, leg. L. Németh (MWM/ ZSM)  ; 5 ♀, valley of Tamea Kosi River, Piguti , 1000m, 7.X.1995, leg. L. Németh (MWM/ ZSM)  ; 3 ♂, 10 ♀, 2200m, 2 km WNW of Muldi ( Murre ), 11.X.1995, 85°54’E, 27°20’N, leg. Hreblay & Bodi (MWM/ ZSM)GoogleMaps  ; 1 ♀, 2200m, Mt. Kalinchok , 2 km WNW Muldi (Murre), 26. VI  .1997, leg. M. Hreblay & K. Csak (MWM/ ZSM); 3 ♂, 2 ♀, 2835m, Mt. Kalinchok , 6 km NNE Muldi (Murre), 27. VI.1997, leg. M. Hreblay & K. Csak, gen. slide No.: ZSM Arct. 2019-505 (male) (prepared by Volynkin) (MWM/ ZSM)  ; 1 ♀, 3100m, Mt. Kalinchok , 8 km NNE of Muldi (Murre), 28. VI  .1997, leg. M. Hreblay & K. Csak (MWM/ ZSM); 4 ♂, 2 ♀, East Nepal, Kanchenjunga Himal , Khambachen, 4150m, 28. VI.1998, leg. Márton Hreblay & Balázs Benedek, gen. slide No.: ZSM Arct. 2019-504 (male) (prepared by Volynkin) (MWM/ ZSM)  ; 2 ♂, 1 ♀, 1850m, Mt. Kalinchok , 8 km E of Barabise, 5.VII.1997, leg. M. Hreblay & K. Csak (MWM/ ZSM)  ; 1 ♀, Mt. Kalinchok area , 5 km NE of Kharidunga, 2950m, 2–3.X.1995, leg. L. Németh (MWM/ ZSM)  ; 2 ♂, East Nepal, Milke Danda, Gursa , 2100m, 22.VIII.2000, leg. Csővári & Hreblay, gen. slide No.: ZSM Arct. 2019-502 (prepared by Volynkin) (Coll. MWM/ ZSM)  ; 1 ♀, the same locality and collectors, 23.VIII.2000 (MWM/ ZSM)  ; 3 ♀, Trisuli valley , 820m, 3 km N Betrawati, 85°11’E, 27°59’N, 25.X.1995, leg. S. Kovács (MWM/ ZSM)GoogleMaps  ; 3 ♀, East Nepal, Deorali Danda, Anpan , 1900m, 18. VI  .1998, leg. Márton Hreblay & Balázs Benedek (MWM/ ZSM); 2 ♀, East Nepal, Deorali Danda , 1 km N of Yamphudin, 2000m, 19. VI  .1998, leg. Márton Hreblay & Balázs Benedek (MWM/ ZSM); 2 ♀, Solu Khumbu Himal, Tragsindha Pass , 3000m, 4.VII.1993, leg. M. Hreblay, G. Csorba (Coll. MWM/ ZSM)  ; 2 ♀, Solu Khumbu Himal, 10 km S of Lukia, Bupsa , 2300m, 3.VII.1993, leg. M. Hreblay, G. Csorba (MWM/ ZSM)  ; 1 ♀, Kathmandu , 1320m, 9–11. VI  .1993, leg. M. Hreblay, G. Csorba (MWM/ ZSM); 1 ♂, Kathmandu , 1235m, 25–27.IX.1994, leg. M. Hreblay & T  . Csővári (MWM/ ZSM)  ; 1 ♀, Phul Chowki, 2000/ 2500m, Kathmandu Valley , J. Plante leg., 6/ 12. VI  .1977 (MWM/ ZSM)  ; 1 ♀, East Nepal, Aruu vall. Num. Nedaugna , 15.IX.–26.X.1980, 750– 1100m, leg. C. Holzschuh (MWM/ ZSM)  . INDIA: 2 ♂, Lumla , Arunachal Pradesh, 10. V  . 2011, 2350m leg. Rahul Joshi (Coll. NZCZSI)  ; 1 ♂, Dentam , Sikkim, 10. V  .2009, leg Rahul Joshi (Coll. NZCZSI)  ; 2 ♂, 1 ♀, W.B., Darjeeling , 2100m, 28.VIII.–1.IX.1988, leg. W. Thomas, gen. slide No.: MWM 17398View Materials (male) (MWM/ ZSM)  ; 1 ♂, 3 ♀, W.B., Darjeeling , Mangpu road, 1800m, 18. VI  .1987, leg. W. Thomas, gen. slide No.: ZSM  Arct. 2020-151 (male) (prepared by Volynkin) (MWM/ ZSM)  ; 2 ♀, W.B., 1900m, Darjeeling, 3 miles vill., 29. VI  .1986, leg. W. Thomas (MWM/ ZSM); 1 ♀, W.B., 2100m, Darjeeling, Ghoom , 8–9.VII.1986, leg. W. Thomas (MWM/ ZSM)  ; BHUTAN: 1 ♂, 1 ♀, Mo Chu River , 24°43’N 89°45’E, 19–20.X.2009, 1500m, leg. Yuri Bezverkhov & Viktor Sinyaev, gen. slide Nos.: AV4229 (male) and AV4230 (female) (Coll. CKC)GoogleMaps  .

Diagnosis. The forewing length is 16.5– 18 mm in males and 19–20 mm in females. The new species is externally similar to its closest relative Z. distorta  , but differs by its somewhat paler forewing. In many cases the reliable identification is possible by the genitalia structures only. The male genital capsule of Z. confusa  sp. nov. differs clearly from that of Z. distorta  by the longer uncus, the shorter juxta, the much broader and slightly curved cucullus (in Z. distorta  , it is straight, narrow and somewhat dilated apically), the more rectangular lobe of the sacculus (that is more rounded in Z. distorta  ), and the broader, more or less bean-shaped and less sclerotized distal saccular process (in Z. distorta  , it is crescent and more heavily sclerotized). The aedeagus of the new species is slightly longer than that of Z. distorta  and bears a longer carina. The aedeagus vesica of Z. confusa  sp. nov. is slightly longer than that of Z. distorta  , tapered apically and curved ventrally, whereas that of Z. distorta  is nearly straight and uniformly wide). The female genitalia of the new species differ from those of Z. distorta  by the conspicuously shorter apophyses posteriores, the somewhat shorter apophyses anteriores, and the broader ductus bursae with convex lateral margins (those are only slightly curved in Z. distorta  ).

Distribution. The species is known from Nepal, Northeastern India (Sikkim, north of West Bengal, Arunachal Pradesh) ( Joshi et al. 2015, as Z. distorta  ), Bhutan and China (Tibet) ( Fang 2000, part). The more eastern records from China ( Fang 2000) are doubtful and need clarification.

Etymology. ‘Confusa’ means ‘confused’ in Latin. The specific epithet refers to its confusion with Z. distorta  .

ZSM

Bavarian State Collection of Zoology

V

Royal British Columbia Museum - Herbarium

VI

Mykotektet, National Veterinary Institute

T

Tavera, Department of Geology and Geophysics

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Lepidoptera

Family

Erebidae

Genus

Zadadra