Praocis (Praocis) tibialis Gay & Solier, 1840

Flores, Gustavo E. & Pizarro-Araya, Jaime, 2012, Systematic revision of the South American genus Praocis Eschscholtz, 1829 (Coleoptera: Tenebrionidae). Part 1: Introduction and subgenus Praocis s. str., Zootaxa 3336, pp. 1-35 : 30-31

Praocis (Praocis) tibialis Gay & Solier, 1840

( Figs. 25 , 30 )

Praocis tibialis Gay & Solier in Solier, 1840: 225 (rev.); Solier, 1851: 193 (rev.); Lacordaire, 1859: 213; Gemminger & Harold, 1870: 1906 (cat.); Philippi, 1887: 733 (cat.); Gebien, 1910: 262 (cat.); Gebien, 1938: 398 (cat.); Blackwelder, 1945: 523 (cat.); Kulzer, 1958: 24 (rev.); Peña, 1966: 430 (cat.); Vidal & Guerrero, 2007: 73; Flores, 2007: 411 (lectot.).

Praocis rufitarsis Gay & Solier in Solier, 1840: 227 (rev.); Solier, 1851: 194 (rev.); Lacordaire, 1859: 213; Gemminger & Harold, 1870: 1906 (cat.); Philippi, 1887: 732 (cat.); Gebien, 1910: 262 (cat.); Gebien, 1938: 398 (cat.); Blackwelder, 1945: 523 (cat.); Kulzer, 1958: 22 (rev.); Peña, 1966: 430 (cat.); Vidal & Guerrero, 2007: 73; Flores, 2007: 411 (syn. & lectot.). Synonymy by Flores, 2007: 411.

Praocis aenipennis Germain, 1855: 400 ; Philippi, 1887: 731 (cat.); Gebien, 1938: 398 (cat.); Kulzer, 1958: 24 (syn.); Peña, 1966: 430 (cat.); Flores, 2007: 411 (type). Synonymy by Kulzer, 1958: 24.

Redescription. Length 6.0–11.0 mm. Body black to dark brown, antennae and legs dark brown. Head. Clypeus and frons with round punctures each with a central seta; antennae reaching 3/4 along lateral margin of pronotum; antennomere 9 longer than wide, with apical tomentose sensory patches arranged in two areas separated dorsally; antennomere 10 as wide as antennomere 11. Thorax. Pronotum widest behind midpoint; disc and lateral quarters with finer golden setae; punctures of disc same size as punctures of elytron; width of anterior margin exceeding half the width of posterior margin; lateral quarters convex, dorsal surface of lateral margin bearing a row of short golden setae; posterior angles right, apex rounded ( Fig. 25 ); prosternum horizontal, with edge on anterior margin broadened below gula; prosternal process produced backwards, not reaching the midpoint of the space between pro– and mesocoxae (as in Fig. 13 ); prosternum and mesosternum with punctures. Elytron lacking grooves, with two or three carinae weakly defined, equidistant between suture and lateral margin ( Fig. 25 ); dorsal surface with short golden setae; pseudopleuron with long setae arising on punctures; epipleuron with short setae arising on protuberances, lacking a row of setae on the edge of anterior quarter. Legs. Ventral surface of meso– and metafemora with a row of setae on posterior edge. Inner surface of meso– and metatibiae with long, fine setae. Metatibiae straight. Male genitalia. Rods of sterna IX “V”-shaped. Basal lamina of tegmen equal width throughout, with base concave. Lateral styles of tegmen with proximal margin ventrally concave, widest at base, with abundant setae on distal third of ventral surface. Median lobe moderate (0.75 <L/T ≤ 1.00), distally broadened, with apical aperture large and apex rounded. Female genitalia. Paraprocts moderate (1.2 P/C ≤ 2.0); baculi of coxite inclined 45°; proctigeral baculus extending proximad much beyond paraproct baculus; apicodorsal lobe of proctiger extending about ½ length of coxite.

Type material. Lectotype of Praocis tibialis : [ tibialis ] [ Praocis / tibialis / Sol./ Illapel] [ P. tibialis G. S. / Sta. Rosa Illapel] [Type] [ Praocis / tibialis Solier / Type de Solier/ Col. Marseul/ C. Girard det. 1975] [Lectotypus/ Praocis tibialis / Gay et Solier, 1840 / Des. G. Flores 2005] ( MNHN). One paralectotype same data as lectotype, plus a label: [ tibialis Sol. 225, 8 Coquimbo] below the label [Type] ( MNHN) .

Note: Flores (2007: 411) designated lectotype.

Lectotype of Praocis rufitarsis : [ rufitarsis ] [ Praocis / rufitarsis / Sol./ Illapel] [Illapel/ Gay] [Type] [ Praocis / rufitarsis Sol. / Type de Solier/ Col. Marseul/ C. Girard det. 1975] [Lectotypus/ Praocis rufitarsis / Gay et Solier, 1840 / Des. G. Flores 2005] ( MNHN) . Three paralectotypes same data as lectotype, one plus a label: [ P. rufitarsis Gay / et Sol. Illapel] and one of the others plus two labels: [ Sa Rosa/ Gay ] and [ P. rufitarsis Sol. / Illapel] ( MNHN) .

Note: Flores (2007: 411) designated lectotype.

Syntype of Praocis aenipennis : [ Praocis / aenipennis / Germain/ Chili] [Chili/ Germain] [ Praocis / aenipennis / P. G (handwritten by Germain )] [ Praocis / tibialis Sol. / (= aenipennis Germ. )/ det. H. Kulzer 1957] ( MNHN) .

Note: This specimen agrees with the description by Germain (1855), and we have no doubt that the label´s handwriting belongs to Germain. Therefore we conclude that this specimen is a syntype of Praocis aenipennis . Kulzer (1958) stated that the type of P. aenipennis is in the MNNC, but currently is not there ( Camousseight 1980 and pers. obs.).

Other material examined. CHILE. Coquimbo Region: Elqui Province: Totoralillo Norte , 21.viii.2009, J. Pizarro-Araya, 1 ( LEULS), Llano La Higuera , 29.ix.1980, L.E. Peña, 1 ( FMNH), El Molle , x.1958, L.E. Peña, 1 ( FMNH), Lengua de Vaca , xi.1978, J. Escobar, 3 ( FMNH). Limarí Province: Talinay , 20.ix.1947, L.E. Peña, 2 ( FMNH), 19.x.2003, J. Pizarro-Araya, 1 ( IADIZA), J. Pizarro-Araya, 1 ( LEULS), 13.x.1976, C. Vivar, 5 ( MNNC), 19.x.2003, J. Pizarro-Araya, 1 ( IADIZA), xi.1978, J. Escobar, 1 ( FMNH), Quebrada Los Maitenes, N Amolanas , 28.ix.1980, L.E. Peña, 1 ( FMNH), Socos , 8.x.1974, L.E. Peña, 2 ( FMNH), 15. xi.2003, 30º 44´S, 71º 31`W, 200 msl, J. Pizarro-Araya 10 ( LEULS), 22. xi.2003, J. Pizarro-Araya 1 ( LEULS), 27.xi.2003, J. Pizarro-Araya 2 ( LEULS). Choapa Province: Hda. Illapel , J. Irarrázabal, 1 ( FMNH), La Higuerilla, Salamanca , 18.ix.2007. A. Gonzalez 3 ( LEULS), 1 ( IADIZA), Huaquilón, San Lorenzo , 24.ix.1982, L.E. Peña, 1 ( FMNH), Puerto Manso , 18.ix.1967, L.E. Peña, 1 ( FMNH), Majada Blanca, Punitaqui , 9.X.2005, J. Pizarro-Araya 3 ( LEULS), S Punitaqui, 7.x.1961, L.E. Peña, 1 ( FMNH), 6 km E Quilimari , 25.xi.1967, L.E. Peña, 2 ( FMNH), 2 ( IADIZA), Choapa, km 272 Panam. Norte , 24.ix.1985, M. Elgueta, 3 ( MNNC), Camino a Mincha , 27.x.1965, L.E. Peña, 1 ( FMNH), Puerto Oscuro , 18–28.ix.1947, L.E. Peña, 1 ( FMNH), 2.ix.1983, A. Roig, 1 ( IADIZA), 23.ix.2004, J. PizarroAraya , 2 ( IADIZA), 23.ix.2004, J. Pizarro-Araya, 1 ( LEULS), Huilmo , 22.ix.1982, L.E. Peña, 1 ( FMNH), Huaquén a Los Molles , 17.xi.1972, L.E. Peña, 1 ( FMNH), Los Molles , 22.x.1961, L.E. Peña, 1 ( FMNH), 2.x.1963, Col. Escuela de Biología , 1 ( FMNH), without more precise data: 15 ( MNNC) .

Geographic distribution. Chile (Coquimbo Region, Coquimbo to Los Molles), in the Coquimbo biogeographic province.

Figures

FIGURE 30. Geographical distribution of five species of Praocis (Praocis): P. (P.) parva, P. (P.) aenea, P. (P.) curta, P. (P.) hirtella, and P. (P.) tibialis.

FIGURES 21 − 26. Habitus in dorsal view. 21. Praocis (Praocis) elliptica; 22. P. (P.) spinolai; 23. P. (P.) medvedevi sp. nov., holotype; 24. P. (P.) aenea; 25. P. (P.) tibialis; 26. P. (P.) parva.

FIGURES 5 − 14. Male and female genitalia and body details of Praocis (Praocis) species. 5. Fore tibia and tarsus in anterior view of Praocis (P.) sanquinolenta. 6 − 9: Male: 6, 7. P. (P.) rufipes, dorsal and ventral views; 8, 9. P. (P.) elliptica, dorsal and ventral views. Abbreviations: bl, basal lamina of tegmen, ls, lateral styles of tegmen, ml, median lobe. 10. Ovipositor (ventral view), spiculum and internal female reproductive tract of P. (P.) hirtella. Abbreviations: c, coxite, o, oviduct, p, paraproct, s, spermatheca, sag, spermathecal accessory gland, sp, spiculum, v, vagina. 11 – 14. Prosternum and mesosternun, lateral aspect. 11. P. (P.) costata; 12. P. (P.) quadrisulcata; 13. P. (P.) elliptica; 14. P. (P.) marginata. Abbreviations: p, prosternum, m, mesosternum. Scale bars: 1 mm.