Our analysis of molecular data on 66 % of the species of Dryopsophus reveals five clades plus outlying species. The first clade has a support value of only 49 % and contains eight species in northern and eastern Australia. Within this group, a well-known species, Dryopsophus caeruleus, is the type species of Pelodryas Günther. A second clade having a support value of 100 % contains four species ranging from Queensland to Victoria in eastern Australia; a member of this group, D. lesueurii, is the type species of Euscelis Fitzinger. Another clade of four species with a support value of 93 % occurs in New Guinea and northern Queensland; no generic name is available for this group. A distinctive clade with a support value of 99 % contains four species of stream-breeding frogs in northern Queensland; one of these, D. nannotis, is the type species of Mosleyia Wells and Wellington. A large group of terrestrial species has been known as Cyclorana Steindachner (type species C. novaehollandiae). In our analysis, this clade has a support value of only 64 %; the clade is widespread in northern Australia and in the interior of the continent. Further, more intense analyses should provide sufficient evidence for the recognition of Cyclorana.
The tadpoles of many species in the mountains New Guinea and in Queensland in northern Australia develop in streams and have enlarged ventral mouths with a LTRF of 2 / 3 ( Tyler 1968; Günther & Richards 2005). Two species in Queensland, Dryopsophus daviesae and D. subglandulosus are unique in not only lacking labial tooth rows but also keratinized beaks ( Anstis 2013).
FIGURE 16. A. Litoria bicolor, KU 179765, AMAX Mining Camp, 300 m, Mitchell Plateau, Western Australia, Australia. W. E. Duellman. B. Litoria nasuta, KU 179853, AMAX Mining Camp, 300 m, Mitchell Plateau, Western Australia, Australia. W. E. Duellman. C. Nyctimystes papua, BPBM 16895, East slope Mt. Simpson, 1440 m, Milne Bay Province, Papua New Guinea. F. Kraus. D. Nyctimystes infrafrenatus, Daintree village area, Queensland, Australia. T. Charlton. E. Dryopsophus lesueurii, KU 179838, Kilcoy Creek, Queensland, Australia. W. E. Duellman. F. Dryopsophus longipes, KU 179923, AMAX Mining Camp, 300 m, Mitchell Plateau, Western Australia, Australia. W. E. Duellman.