Shoreeae Miq., Fl. Ned. Ind.

Cvetković, Tijana, Hinsinger, Damien D., Thomas, Daniel C., Wieringa, Jan J., Velautham, Elango & Strijk, Joeri S., 2022, Phylogenomics and a revised tribal classification of subfamily Dipterocarpoideae (Dipterocarpaceae), TAXON 71 (1), pp. 85-102 : 92-93

Tribe Shoreeae Miq., Fl. Ned. Ind. 1(2): 503. 1859

– Type: Shorea Roxb. ex C.F.Gaertn.

= [unranked] Hopeae Korth., Verh. Nat. Gesch. Ned. Bezitt., Bot.: 55. 1840.

– “Parashoreeae” Takht., Flowering Pl.: 269. 2009, nom. nud. – Based on Parashorea Kurz.

Emergent or canopy, understorey trees; resin canals in tangential bands; thickened sepal base; fruit sepals imbricate at the incrassate-cupped base of the ripe fruit; 3 strata in pollen exine: absent tilioid structure of exine; T and Y columellae shape-type; always grouped vessels with cellular divisions; radial canal formation; 2 or 3 incrassate bases of sepals (and accrescent sepals) in fruits; free bases of fruit sepals; frigid pericarp tissue; circular fruit equatorial section; embryo cotyledons “coveringpiled”; inferior or median-inferior hypocotyl; bilobed seedling cotyledons; 4 root-xylem poles; uni- to tri-lacunar cotyledonary vascular bundles; stomatal types in first leaves paracytic, or para-cyclocytic, or anomo-cyclocytic; elongate stomata, sunken in the epiderm; chromosome number n = 7 (summarized in Appanah & Turnbull, 1998). From India and Sri Lanka to Malesia.

Eight genera: Anthoshorea Pierre (synonym of Shorea ), Doona Thwaites (synonym of Shorea ), Hopea Roxb. , Neobalanocarpus P.S.Ashton , Parashorea Kurz, Richetioides F.Heim (synonym of Shorea ), Rubroshorea (synonym of Shorea ), Shorea Roxb. ex C.F.Gaertn.

Notes: Hopeae, described by Korthals (1839 –1842), would have had priority over Shoreeae but was described without explicit rank, which is mandatory before 1887 (Art. 37.1–37.3 of the ICN, Turland & al., 2018).

Shoreeae has also been spelled as ‘Shoreae’. Given this name has a non-Latin base (named after Sir John Shore), but has been Latinized, where the stem is the single syllable Shore, it seems proper to follow the original spelling of Miquel, who also used ‘Shoreeae’. In case we consider this name to have alternative possible genitives, Art. 18.1 (via Art. 17.1) even obliges us to do so. In botanical Latin, double ee’ s are often avoided, but many recent tribal names based on generic names ending on -ea have also regularly been formed including the double ee. Since Parashorea is derived from Shorea , a tribe named after it should be spelled with the double ee as well, but so far this name lacks a formal description and is not in alignment with our classification superfluous.