Ectatops riedeli, Stehlík & Jindra, 2008

Stehlík, Jaroslav L. & Jindra, Zdeněk, 2008, New taxa of the Largidae and Pyrrhocoridae (Hemiptera: Heteroptera) from the Oriental Region, Acta Entomologica Musei Nationalis Pragae 48 (2), pp. 611-648 : 640-643

Ectatops riedeli sp. nov.

(Figs. 42, 44, 48) Type material. HOLOTYPE: J, INDONESIA: SULAWESI: SULAWESI SELATAN PROVINCE: ca. 15 km W of Palopo, 18.- 19.viii.1990, A. Riedel lgt. (ZSMC). PARATYPES: The same data, 1 J 1 ♀ (ZSMC).

Description. Colouration (Fig. 42). Black on head (except orange spot on vertex), antennomeres 1-3 (except bases), antennomere 4 narrowly at base and slightly more than its apical half, scutellum (except of apex), membrane (except of base), and entire ventral side of body.

Figs. 40-43. Habitus. 40 – Dysdercus (Paradysdercus) transversalis castaneus subsp. nov. (holotype, ♀, 15.10

mm); 41 – Euscopus tristis sp. nov. (holotype, J, 7.40 mm); 42 – Ectatops riedeli sp. nov. (holotype,J, 8.99 mm);

43 – E. schoenitzeri sp. nov. (holotype, J, 8.27 mm). Photo: L. Dembický.

Pronotal lobe, clavus, and corium brown (somewhat darkened towards apex, veins of corium also darkened); in females all of them somewhat darker with costal margin laterally pale at level of claval comissure. Narrow basal ring on antennomeres 1-3, ring in basal half of antennomere 4, lateral margin of pronotum, and sometimes also pronotal collar and pronotal epipleuron pale brown. Apices of scutellum and corium white yellow. Ventral laterotergites black, dorsal laterotergites dark brown with outer edge somewhat paler. Profemora black, meso- and metafemora basally very pale brown, then gradually blackend towards apex; narrow rings on apices of femora and bases of tibiae and tarsomere 1 (except of apex) whitish or yellowish. Rather wide rings on bases of tibiae, narrow rings at their apices, apex of tarsomere 1 and entire tarsomeres 2 and 3 blackish.

Structure. Body relatively small. Eye sockets strong, produced laterally and distinctly raised. Anterior part of head shorter, more strongly sloping.Antennae slender; antennomere 1 slightly thickened towards apex; antennomere 2 very slightly thickened apically; antennomere 3 gradually widening from base, apically wider than antennomere 2; antennomere 4 widest. Callar lobe regularlly gibbous; lateral pronotal margin at level of callar lobe rounded, at level of callar lobe concave, at level of pronotal lobe more distinctly diverging sideways.

Pygophore ( Fig. 44 ). Ventral part of ventral wall strongly gibbous, emarginated by submedially skewed furrows pointing ventrally; upper part of ventral wall medially rather widely depressed. Ventral rim submedially (on both sides) with whitish, rather wide process skewed inwards genital chamber, covered by long pale hairs; dorsal margin of the process rounded, its posterior corner (directed into genital chamber) slightly pointed. Lower margin of lateral rim infolding strongly elevated and nearly semicircularly enfolding two dark, back-to-back standing processes, whoes upper facet is medially concave. Body of paramere thickened, distal part thinner and narrowed before apex; edge of apex medially slightly concave and produced laterally into small spines. Inner side of paramere with another dark comb ( Fig. 48 ). Distal part of vertical process nearly rectangular, margins slightly concave, apex rounded.

Female outer genitalia much wider than high. Both sides of valvifer parallel near base, then arcuatelly diverging, its upper margin only slightly raised dorsally. Laterotergite VIII triangular; laterotergite IX large, triangularly narrowed towards centre of genitalia, flat, only its margins narrowly ridged. Anal tube slit-like. Upper margin of valvifer and laterotergite IX pale pubescent.

Punctation on pronotal lobe, clavus, and corium prominent, black, of similar size. Mesoscutum punctate; punctures in apical part of scutellum hardly visible on black colouration.

Measurements (all in mm). Males (holotype first, paratype second). Body length 8.99/8.96; head: length 1.57/1.46, width (including eyes) 2.32/2.32, interocular width 1.24/1.30; lengths of antennomeres: 1 – 1.73/1.40, 2 – 1.35/1.40, 3 – 1.16/1.16, 4 – 1.43/1.43; pronotum: length 1.81/1.89, width 2.27/3.02; scutellum: length 1.13/1.19, width 1.57/1.57; corium: length 3.89/4.19, width 1.51/1.67.

Female (paratype). Body length 9.94; head: length 1.51, width (including eyes) 2.52, interocular width 1.35; lengths of antennomeres: 1 – 1.94, 2 – 1.62, 3 – 1.35, 4 – 1.57; pronotum: length 2.27, width 3.48; scutellum: length 1.35, width 1.94; corium: length 4.97, width 2.00.

Differential diagnosis. Ectatops riedeli sp. nov. (Fig. 42) is very similar both in colouration and external morphological characters to E. sulawesiensis Stehlík & Jindra, 2006 (from northern Sulawesi) and E. subjectus Walker, 1873 (from central Sulawesi, Butung Island). However, all three species are easily recognizable by the structure of the ventral rim on the pygophore and the paramere. Ectatops sulawesiensis is larger (body length 9.45-10.0 mm in males); ventral rim on its pygophore submedially with one stout, dark process on each side, slightly inclined medially and inwards the genital chamber, on outer side slightly concave, apically rounded, on inner side of apex somewhat spinous; dorsal part of ventral wall and ventral rim medially depressed (as in E. riedeli sp. nov.); vertical process more slender, apically somewhat widened and rounded (as in E. riedeli sp. nov.) ( Fig. 46 ); body of paramere basally wide, before end of this part with a small denticle and longer sensory setae above the denticle; apical part of paramere long, slender, somewhat curved in two thirds, apex horizontally truncated with minute denticles directed sideways ( Fig. 50 ).

Ectatops subjectus is of the same size as E. riedeli sp. nov. but well recognizable by the following characters: ventral rim submedially with one pale stylus on each side, somewhat thickened towards apex and bearing prominent pale pilosity (very different from the wide, nearly quadrangular process of E. riedeli sp. nov.); ventral rim widely incised (not depressed as in E. riedeli sp. nov. and E. sulawesiensis ), incision flat and horizontal; vertical process wider than in E. riedeli sp. nov. and E. sulawesiensis , parallel-sided, deeply triangularly incised ( Fig. 45 ); body of paramere wider, on outer side shorter, on inner side slightly longer, with longer dark comb, above the comb incised, with short pale hairs; apical part wider, narrowed before apex, curved; apex black with small spicula on one side and the other side projected into long, elevated spicula ( Fig. 49 ).

Etymology. We dedicate this species to Alexander Riedel (Karlsruhe, Germany), whose field trips to the Oriental Region and New Guinea resulted in the discovery of many undescribed species.

Distribution. Indonesia, south-western Sulawesi.

Figures

Figs. 44-50. 44-47 – pygophore (posterior view): 44 –Ectatops riedeli sp. nov.; 45 – E. subjectus Walker, 1873; 46 – E. sulawesiensis Stehlík & Jindra, 2006; 47 – E. schoenitzeri sp. nov. 48-50 – parameres: 48 – E. riedeli sp. nov.; 49 – E. subjectus (two different views of apical portion); 50 – E. sulawesiensis.