Ephutomma himalayana Lelej and Ullah

Lelej, Arkady S., Ullah, Mishkat & Mahmood, Khalid, 2007, Additions to the knowledge of the Mutillidae (Hymenoptera) of Pakistan, Zootaxa 1444, pp. 53-60 : 58

Ephutomma himalayana Lelej and Ullah , sp. nov.

( Figs 16–25 )

Diagnosis. The male of Ephutomma himalayana is most similar to that of E. kerzhneri (Lelej 1977) [holotype is examined, deposited in ZIN] in the key to the Palaearctic species ( Lelej 1985). It can be distinguished from the latter species in having an additional subapical tooth on the ventral mandibular margin (lacking in E. kerzhneri ), in having a subbasal ventral mandibular tooth that is unidentate apically (bidentate in E. kerzhneri ), and in having infuscated wings (hyaline with infuscated apex in E. kerzhneri ).

Description. MALE. Body length 5.4–9.6 mm (holotype – 7.2 mm). Head width 1.2X head height. Ocelli small, POD: OOD 0.5–0.6X. Clypeus with anterior shiny depression bordered by arcuate carina which ends anteriorly in two tubercles. Occipital carina weak. Hypostomal carina weakly dilated. Mandible dilated, with inner tooth and additional subapical ventral tooth, bidentate apically, excised beneath, subbasal ventral tooth with one apex ( Figs 18–21 ). Relation of pedicel and three first flagellomeres 0.5: 1.0: 2.0: 2.0. Frons without process between antennal tubercles. Ocellar area weakly elevated. Frons, vertex, pronotum, mesoscutum, scutellum, and mesopleura densely punctate. Mesoscutum with complete notauli and short parapsidal furrows. Parascutal carinae well developed. Propodeum reticulate with median elongated and two rounded cells on dorsum. Mesopleura beneath without precoxal tubercle. Posterior coxae carinate inside, carina not dentate apically. Tegulae punctate, disc and posterior margin glabrous. Forewing venation as in Fig.16 . S1 carinate beneath. Metasomal segment 2 densely punctate, punctures sparser on tergal disc. T2 with long lateral felt line. S2 without any trace of felt line. T3–T6 densely punctate. T7 densely punctate, apically with median glabrous area. Genitalia laterally, dorsally, and ventrally as in Figs. 22–25 .

Body, legs, and antennae black; pronotum, mesoscutum, scutellum, tegulae, metanotum, upper part of mesopleura, and upper part of metapleura red. Wings infuscated. Mandibles preapically piceous. Fore tibial spurs yellowish, mid- and hind tibial spurs whitish. Lower frons, genae, antennal tubercles, scape, tibiae, and tarsi covered with dense, recumbent short and scattered, long, erect whitish pubescence. Upper frons, vertex, pronotum, scutellum, tegulae, metanotum medially, propodeum, T1–T8, S1–S8, and femorae covered with sparse whitish setae. Mesoscutum with sparse black setae. Scutellum and metanotum laterally covered with dense recumbent silvery micropubescence. T1–T3 with apical fringe of dense yellowish setae. S2–S5 with apical fringes of sparse whitish setae.

FEMALE. Unknown.

Type material. Holotype ♂, Nepal, Chitwan National Park, Sauraha, 27°34'N, 84°29'E, 200 m, 31.V.1997, M. Hauser [Oberoesterreichisches Landesmuseum Linz, Austria]. Paratypes. Nepal: the same label as holotype, 1 ♂ [ IBSS]; the same place, 31.V–4.VI.1997, M. Hauser, 3 ♂ [col. Schmid-Egger, IBSS]. Pakistan: Margalla Hills National Park, 1, 13, 23.VI, 1, 12.VII, 8.IX.2005, MT, H. Khurram & M. Fida, 6 ♂ [ PMNH, IBSS]; 25.VIII.2006, MT, H. Khurram & M. Fida, 1 ♂ [ PMNH].

Distribution. Pakistan, Nepal.

Etymology. The specific name is a Latin adjective derived from the Himalaya Mountain range, referring to the region where the species has been found.

Figures

FIGURES 16 – 25. Ephutomma himalayana sp. nov., male, paratype. 16. Forewing; 17. Pedicel and flagellomeres 1 – 3, ventral view; 18 – 21. Mandible (18 – outer view, 19 – inner view, 20 – ventral view, 21 – dorsal view); 22 – 25. Genitalia (22 – lateral view, 23 – ventral view, 24 – dorsal view, 25 – volsella).