Lepanus reidi Gunter & Weir, 2019

Gunter, Nicole L. & Weir, Thomas A., 2019, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups, Zootaxa 4695 (3), pp. 201-252 : 231-234

Lepanus reidi Gunter & Weir , new species

( Figs 8B, F, H ; 9C ; 10C ; 11C .)

Type series. Holotype: ♂ “QLD: 17.430°S 145.490°E Crater NP, 18-19 Sep 2010, G. Monteith Dung trap, RF, 980m 34584” (= Mout Hypipamee National Park ) / “ ANIC Database No. 25 059231” / “ ANIC image” / “ Lepanus NQ5 det TA Weir 2010 ”, in ANIC (17°25’47”S, 145°29’24”E) . Paratypes (25 ♂, 35 ♀, 5 sex undetermined): Crater National Park , 980 m, dung trap 34584, 17.430°S, 145.490°E 18–19.ix.2010, G.B. Monteith (25- 060883 3 specimens in ANIC also as COL1152 ♀, COL1153 ♀, COL1154 ♀) (17°25’47”S, 145°29’24”E) ; Mount Hartley , 2.5 km SW, 15°47’49”S, 145°19’34”E, 24.iv.1982, ( T 54260 in QM 1 specimen ♀) ; Mount Finnigan S1, 30 km S of Cooktown, [15°49’0”S, 145°17’0”E], 400 m, 2.vii.1982, S. & J. Peck (25-059106 1 specimen ♂ 1 specimen ♀ in ANIC) ; Alexandra Range , 16°14’42”S, 145°26’10”E, 200 m, 20–21.ix.2010, G.B. Monteith (25- 060882 1 specimen in ANIC also as COL1151 ♀) ; Mount Sorrow track, 16°4’37”S, 145°27’58”E, 30 m, 28–30.v.2012, G.B. Mon- teith (25-057612 1 specimen, 25-057617 1 specimen ♀ also as COL 2448in ANIC) ; Emmagen Creek , 16°2’24”S, 145°27’32”E, 10 m, 29–30.v.2012, G.B. Monteith (25-057618 2 specimens ♀ also as COL 2450 in ANIC) ; 1 km WNW Cape Tribulation , [16°4’0”S, 145°28’0”E], 10 m, x.1996, J. Seymour (25-059136 1 specimen ♀ in ANIC) ; Cape Tribulation , [16°5’0”S, 145°27’0”E], 20 m, 9.v.1997, C. Reid (25-059134 1 specimen ♀, 25-059135 1 speci- men ♀ in ANIC) ; Cape Tribulation , 40 km N of Daintree, [16°5’0”S, 145°29’0”E], 10 m, 12–15.vii.1982, S. & J. Peck (25-059105 3 specimens ♂, 2 specimens ♀ in ANIC) ; Oliver Creek , 16°8’22”S, 145°26’24”E, 4–7.vi.1974, ( T60127 1 specimen ♂ in QM) ; Alexandra Range , N slope, 16°14’31”S, 145°25’48”E, 140 m, 28–30.v.2012, G.B. Monteith (25-057616 1 specimen ♂ also as COL 2449 in ANIC) ; Windsor Tableland ANZSES Site 9, 16°16’55”S, 145°3’56”E, 27.xii.1988 – 8.i.1989, ( T54257 1 specimen ♀ in QM) ; Mossman Bluff Track 5–10 km W Mossman, Site 1, 16°28’19”S, 145°19’37”E, 20.xii.1989 – 15.i.1990, ( T54269 – T54270 1 specimen ♀, 1 specimen sex undeter- mined in QM) ; Mossman Bluff Track 5–10km W Mossman, Site 1, 16°28’19”S, 145°19’37”E, 1.i.1989 – 16.i.1989, ( T54271 – T54279 7 specimens ♂, 2 specimens ♀ in QM) ; Mossman Gorge, 16°28’26”S, 145°19’54”E, 1.xi.1991 – 22.vii.1992, ( T54258 – T54259 1 specimens ♂, 1 specimens ♀ in QM) ; Black Mountain , lower slopes, 16°39’52”S, 145°30’4”E, 2.xii.1997 – 6.ii.1998, ( T60126 1 specimen ♀ in QM) ; GS1 Mount Haig , [17°6’0”S, 145°36’0”E], 1150 m, 1.xii.1994 – 3.i.1995, P. Zborowski (25-054103 1 specimen ♂, 2 specimen ♀ in ANIC) ; Massey Range , 2nd camp, 17°13’51”S, 145°48’13”E, 11–12.x.1991, ( T54256 1 specimen ♀ in QM) ; Bartle Frere , West Base, 17°22’47”S, 145°45’51”E, 25.xi.1994 – 10.i.1995, ( T54261 – T 54264 in QM 1 specimen ♂, 3 specimen ♀) ; Bartle Frere , West Base, 17°22’47”S, 145°45’51”E, 10.i–31.iii.1995, ( T54265 – T54266 2 specimens ♀ in QM) ; Bartle Frere , West Base, 17°22’47”S, 145°45’51”E, 7.iii–15.v.1995, ( T54267 1 specimen sex undetermined in QM) ; BS3 Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 1.xii.1994 – 3.i.1995, P. Zborowski (25-054101 3 specimens ♂, 3 speci- mens ♀ in ANIC) ; BS3 Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 3.x–2.xi.1995, L. Umback (25-054105 1 specimen ♂ in ANIC) ; Mount Fisher site BS2, [17°33’0”S, 145°32’0”E], 1150 m, 10.xii.1994 – 3.i.1995, P. Zborows- ki (25-054104 1 specimen ♀ in ANIC) ; Maalan Road , 2 km S Palmerston Highway, 17°35’56”S, 145°41’58”E, 7.iii–15.v.1995, ( T54252 – T54253 1 specimen ♂, 1 specimen ♀ in QM) ; Maalan Road , 2 km S Palmerston Highway, 17°35’56”S, 145°41’58”E, collected 26.xi.1994 – 10.i.1995, ( T54254 – T54255 1 specimen ♂, 1 specimen sex unde- termined in QM) ; Bellenden Ker Range , 0.5 km S Cable Tower 7, [17°16’0”S, 145°55’0”E], 500 m, 25–31.x.1981, Earthwatch, Queesland Museum, (25-059108 1 specimen ♂ in ANIC) ; Bellenden Ker Range , Cableway Base Sta- tion, [17°16’0”S, 145°55’0”E], 100 m, 17.x–9.xi.1981, Earthwatch, Queensland Museum, (25-059110 1 specimen sex undetermined in ANIC) ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 1.xii.1983 – 9.i.1984, R.I. Storey & Brown (25-059130 1 specimen ♂, 1 specimen ♀ in CMNC) ; Mount Father Clancy , 800 m, 17°35’58”S, 145°38’4”E, 6–14.xii.1988, ( T54219 1 specimen ♀ in QM) ; Palmerston National Park , E margin, 17°36’40”S, 145°45’48”E, 2.xi.1995 – 10.xii.1995, ( T54210 1 specimen ♂ in QM) .

Other material examined. 427 specimens from almost 50 localities. Mount Boolbun South , (camp in QM) , 15°57’10”S, 145°8’’23”E, 5.xi.1995 – 11.i.1996, ( T54191 – T 54195 in QM) ; Mount Boolbun South , (camp in QM) , 15°57’10”S, 145°8’’23”E, 5.xi.1995 ( T 54196 in QM); Mount Boolbun South , (camp in QM), 15°57’10”S, 145°8’’23”E, 4–6.xi.1995, ( T54197 – T 54201 in QM) ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 14.i– 10.ii.1998, ( T 73143 in QM) ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 10.ii.1998 – 14.iii.1998, ( T 85650 in QM) ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 14.iii– 6.v.1998, ( T88788 – T88790, T88793 – T 88794 in QM) ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 10.ix–26.x.1998, ( T88791 – T 88792 in QM) ; Cape Tribulation transect Site 1, 16°4’12”S, 145°27’54”E, 23.ix–7.x.1982, ( T 54165 in QM) ; Cape Tribulation transect Site 2, 16°4’28”S, 145°27’33”E, 20–23.iv.1983, ( T54162 – T 54164 in QM) ; Cape Tribulation transect Site 2, 16°4’28”S, 145°27’33”E, 23.ix –7.x.1982, ( T 54166 in QM) ; Cape Tribulation transect Site 5, 16°4’41”S, 145°27’13”E, 20–23.iv.1983, ( T54146 – T 54160 in QM) ; Cape Tribulation transect Site 5, 16°4’41”S, 145°27’13”E, 5–9.i.1983, ( T 54161 in QM) ; Cape Tribulation transect Site 5, 16°4’41”S, 145°27’13”E, 23.ix –7.x.1982, ( T54185 - T 54186 in QM) ; Cape Tribulation transect Site 5A, 16°4’41”S, 145°27’0”E, 20–23.iv.1983, ( T54182 – T 54184 in QM) ; Cape Tribulation transect Site 6, 16°4’41”S, 145°26’39”E, 23.ix-7.x.1982, ( T54167 – T 54177 in QM) ; Cape Tribulation transect Site 6, 16°4’41”S, 145°26’39”E, 5–9.i.1983, ( T54178 – T 54180 in QM) ; Cape Tribulation tran- sect Site 6, 16°4’41”S, 145°26’39”E, 20–23.iv.1983, ( T 54181 in QM) ; Thompson Creek, ( Grove ), 16°6’’31”S, 145°26’25”E, 10.i–12.ii.1998, ( T73153 – T 73154 in QM) ; Thompson Creek, (Grove), 16°6’’31”S, 145°26’25”E, 15.iii.1998 – 7.v.1998, ( T 88806 in QM) ; Oliver Creek, 16°8’’22”S, 145°26’24”E, 4–7.vi.1974, ( T 49882 in QM) ; Cooper Creek, ( Grove ), 16°9’10”S, 145°24’19”E, 10.ii–16.iii.1998, ( T85646 – T 88801 in QM) ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 8.i–9.ii.1998, ( T73155 – T 73160 in QM) ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 9.ii–13.iii.1998, ( T85640 – T 85649 in QM) ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 13.iii–8.v.1998, ( T88807 – T 88812 in QM) ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 2– 10.xii.2012, T.A. Weir & G.B. Monteith, (25-057656 1 specimen, 25-057658 4 specimens, 25-057662 3 specimens in ANIC) ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 6–9.xii.2012, G.B. Monteith (25-057652 2 specimens, 25-057654 x 1 in ANIC) ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 9.xii.2012, T.A. Weir (25-057665 3 specimens in ANIC) ; Mangrove Road, ( Grove ), 16°12’42”S, 145°25’18”E, 8.i-9.ii.1998, ( T73144 – T 73152 in QM) ; Mangrove Road, ( Grove ), 16°12’42”S, 145°25’18”E, 9.ii–13.iii.1998, ( T 85642 in QM) ; Mangrove Road, ( Grove ), 16°12’42”S, 145°25’18”E, 9.ix–25.x.1998, ( T88813 – T 88815 in QM) ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 13.i–11.ii.1998, ( T73161 – T 73167 in QM) ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 11.ii–14.iii.1998, ( T85643, T 85648 in QM) ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 14.iii –8.v.1998, ( T88802 – T 88805 in QM) ; Fairy Creek, ( Grove ), 16°14’16”S, 145°24’57”E, 11.ii– 13.iii.1998, ( T 85647 in QM) ; Fairy Creek, ( Grove ), 16°14’16”S, 145°24’57”E, 13.iii–8.v.1998, ( T 88787 in QM) ; Buchanan Creek , (Grove in QM) , 16°14’39”S, 145°25’54”E, 5.i–11.ii.1998, ( T73141 – T 73142 in QM) ; Buchanan Creek , (Grove in QM) , 16°14’39”S, 145°25’54”E, 12.ix–23.x.1998, ( T88795, T 88797 in QM) ; Buchanan Creek , (Grove in QM) , 16°14’39”S, 145°25’54”E, 12.iii–8.v.1998, ( T88796, T88798 – T 88799 in QM) ; Windsor Tableland via Mount Carbine , [16°15’0”S, 145°2’0”E], 25.xi.1976, R.I. Storey (25-059124 8 specimens in QDPC) ; Windsor Tableland ANZSES Site 9, 16°16’55”S, 145°3’56”E, 27.xii.1988 – 8.i.1989, ( T 54188 in QM) ; Julatten (general), [16°37’0”S, 145°20’0”E], 21–30.xi.1987, A. Walford-Huggins (25-059114 5 specimens in ANIC) ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 10.xii.1984 – 15.i.1985, R.I. Storey & H.K. Halfpapp, (25-059118 6 specimens in ANIC) ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 15.i.1985 – 20.ii.1985, R.I. Storey & H.K. Halfpapp, (25- 032604 1 specimen in QDPC, 25-059117 5 specimens in QDPC) ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 20.ii–20.iii.1985, R.I. Storey & H.K. Halfpapp, (25-059120 1 specimen in QDPC) ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 6.xi–10.xii.1984, R.I. Storey & H.K. Halfpapp, (25-059119 4 specimens in QDPC) ; North Bell Peak summit, 17°5’19”S, 145°52’44”E, 15–16.ix.1981, ( T 54268 in QM) ; Bellenden Ker Range, Cableway Base Station , [17°16’0”S, 145°55’0”E], 100 m, 17.x–9.xi.1981, Earthwatch, Queensland Museum, (25-059109 1 specimen in ANIC) ; Graham Range , summit, 17°16’24”S, 145°57’58”E, 1.xi–8.xii.1995, ( T 54240 in QM) ; Graham Range , 350 m, 17°16’37”S, 145°57’48”E, 1.xi–8.xii.1995, ( T54237 – T 54239 in QM) ; Wongabel State Forest , 6 km S of Atherton, [17°18’0”S, 145°31’0”E], 1.v–21.vi.1984, R.I. Storey & Brown (25-059126 1 specimen in ANIC) ; Topaz, Pei Road , (Frith house in QM) , 17°24’23”S, 145°41’5”E, 6.xii.1993 – 25.ii.1994, ( T54242 – T 54244 in QM) ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 1.xii.1983 – 9.i.1984, R.I. Storey & Brown (25- 032603 1 specimen, 25-059111 4specimens in QDPC) ; Hugh Nelson Range , 21km S of Atherton, [17°25’0”S, 145°28’0”E], 1. v–21.vi.1984, R.I. Storey & Brown (25-059129 1 specimen in QDPC) ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 10.ii–13.iii.1984, R.I. Storey & Brown (25-059127 2 specimens in QDPC) ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 12.x–5.xi.1983, R.I. Storey & Brown (25-059131 1 specimen in QDPC) ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 13.iii–1.v.1984, R.I. Storey & Brown (25-059128 3 specimens in QDPC) ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 5.xi–1.xii.1983, R.I. Storey & Brown (25-059132 1 specimen in QCPC) ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 9.i–10.ii.1984, R.I. Storey & Brown (25-059133 1 specimen in QDPC) ; The Crater National Park , [17°25’0”S, 145°29’0”E], 16–18.iv.1973, R.I. Storey (25-058484 1 specimen in QM) ; Topaz, Hughes Road , (Breeden house in QM) , 17°26’4”S, 145°42’2”E, 6.xii.1993 – 25.ii.1994, ( T 54241 in QM) ; Millaa Millaa Falls , 17°29’53”S, 145°36’32”E, 6–9.ii.1999, ( T76200 – T 76201 in QM) ; Millaa Millaa Falls (below), [17°29’0”S, 145°36’0”E], 760 m, 18–20.iv.1999, C. & I. Reid (25-059112 1 specimen in ANIC) ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 10.iv–4.v.1990, R.I. Storey & H.K. Halfpapp, (25- 032602 1 specimen in QDPC) ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 13.iii–10.iv.1990, R.I. Storey & H.K. Halfpapp, (25- 032601 3 specimens in QDPC) ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 4.i–7.ii.1990, R.I. Storey & H.K. Halfpapp, (25- 032599 8 specimens in QDPC) ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 7.ii–13.iii.1990, R.I. Storey & H.K. Halfpapp, (25- 032600 4 specimens in QDPC) ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 7.iii–15.v.1995, ( T54246 – T 54247 in QM) ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 10.i–31.iii.1995, ( T54248 – T 54250 in QM) ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 1.xii–25.ii.1994, ( T 54245 in QM) ; Mount Fisher site BS2, [17°33’0”S, 145°32’0”E], 1150 m, 4.ix–3.x.1995, L. Umback (25-055628 1 specimen in ANIC) ; Maalan SF, Ravenshoe Road, 17°35’23”S, 145°35’40”E, 7.iii–15.v.1995, ( T54202 – T 54204 in QM) ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 10.i–31.iii.1995, ( T 54205 in QM) ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 10.i–31.iii.1995, ( T54206 – T 54208 in QM) ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 25.xi.1994 – 10.i.1995, ( T 54209 in QM) ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 6–9.ii.1999, ( T76197 – T 76199 in QM) ; Mount Father Clancy , 900 m, 17°35’45”S, 145°38’4”E, 4.v.1983, T 54187 in QM ); Rosina Creek , 14 km SE Millaa Millaa , [17°36’0”S, 145°43’0”E], 720 m, 24.vi–2.viii.1982, S. & J. Peck (25-059107 1 specimen in ANIC) ; BS3 Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 1.xii.1994 – 3.i.1995, P. Zborowski 25-054102 5 specimens in ANIC ); BS3 Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 3.i–5.ii.1995, P. Zborowski (25-054106 1 specimen in ANIC) ; Plamerston National Park , [17°37’0”S, 145°47’0”E], 22–24.iv.1979, R.I. Storey (25-058483 1 specimen in QM) ; South Johnstone Forestry Camp , 5 km W, 17°38’12”S, 145°40’15”E, 1.xii.1993 – 25.ii.1994, ( T54211 – T 54214 in QM) ; Charmillin Creek crossing, 17°42’3”S, 145°31’21”E, 8.xii.1989 – 5.i.1990, ( T54215 – T 54218 in QM) ; Ravenshoe State Forest, Tully Falls Road, [17°43’0”S, 145°33’0”E], 10.x–15.xi.1987, A. Walford-Huggins (25-059113 1 in ANIC) ; Ravenshoe State Forest, Tully Falls Road, [17°43’0”S, 145°33’0”E], 15.xi–12.xii.1987, A. Walford-Huggins (25- 059115 1 in ANIC) ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 3.iii– 20.iv.1988, R.I. Storey & Dickinson, (25-059125 1 in QDPC) ; Tully Falls , 17°47’0”S, 145°33’27”E, 8.xii. 198 9– 5.i.1990, ( T 54220 in QM) ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 1.x–5.xi.1987, R.I. Storey & Dickinson, (25-059116 2 specimens in QDPC) ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 5.i–7.ii.1988, R.I. Storey & Dickinson, (25-059122 2 specimens in QDPC) ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 5.xi–7.xii.1988, R.I. Storey & Dickinson, (25-059121 3 specimens in QDPC) ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 7.xii.1987 – 7.i.1988, R.I. Storey & Dickinson, (25-059123 2 specimens in QDPC) ; Boulder Creek track, 200–600 m, 17°51’2”S, 145°54’29”E, 24–27.x.1983, ( T54221 – T 54236 in QM) .

Description. Holotype uniformly light brown. Others ranging from light brown to dark brown almost black. Antennal clubs grey.

Total length: 2.4–2.8 mm. Holotype measurements: total length 2.6 mm, elytral width 2.0 mm.

Male. Head. Surface very finely punctate, often appearing almost impunctate, basal carina absent. Clypeal edge normal, without projection lateral to teeth. Genal angle weakly defined. Dorsal part of the eyes narrow, separated by an interocular space approximately 20 times the eye width (40: 2); eye canthus nearly dividing eye. Head 1.3 x wider than long (48:36).

Prothorax: Pronotal surface with very fine punctation. Basal angles distinct. Punctures along the basal margin absent. Pronotum 1.8 x wider than long (41:23).

Elytra: Surface smooth with obsolete striae; intervals virtually impunctate. Striae 6 and 7 extending to elytral base. Elytra 1.1 x wider than long (51:45).

Legs: Protibiae usually with 3 teeth on outer edge, sometimes with only 2 teeth, front edge deeply angulate. Apical digit long and parallel sided with a rounded apex with a very small comb of stout setae, and bearing a short spur on the outer apical angle. Metatibiae with a notch on the inner edge.

Abdomen: Pygidium simple, entirely smooth, virtually impunctate and very slightly convex. Abdominal ventrites smooth, shiny and without reticulation. Ventrite 6 very finely punctate, almost impunctate.

Pterothorax: Medial lobe of metaventrite very finely punctate and very narrowly margined between the mesocoxae. Lateral lobe of metaventrite with inner border beside mesocoxae anteriorly curving to meet lateral border. Mesoventrite notched anteriorly. Mesoventrite smooth. Metanepisternum finely reticulate.

Aedeagus: Small tufts of setae projecting from apices of both right and left parameres, left paramere pointed.

Female. As per L. monteithi species group. Metatibiae without notch on inner edge.

Etymology. Named in honour of Chris Reid, at the Australian Museum, for his invaluable contribution to the systematics of Australian beetles including the dung beetle genera Temnoplectron and Coptodactyla .

Distribution. Known from the rainforests of the Wet Tropics bioregion from Mount Boolbun south to Tully Falls National Park ( Fig. 12B ).

Comments. Lepanus reidi is known from almost 500 specimens, the majority of which have been collected in flight intercept traps or baited pitfall traps. This species appears to be a generalist and label data indicate this species has been collected in earthworm, cricket, or snail baited flight intercept traps, diurnally feeding on bird faeces, and from mushroom or dung baited pitfall traps (human or macropod faeces). Ebert et al. (2019) also collected L. reidi new species from a pitfall trap baited with vertebrate carrion. A small number of specimens have been collected in malaise and window traps.

This species has been referred to under the informal nomenclature sensu Yeates et al. (2011) as Lepanus NQ5 (Gunter et al. 2019). The specimen sequenced in this study was collected from the Mount Hypipamee National Park [formerly The Crater National Park] in the same trap as the holotype and is designated as a paratype here (ANIC 25- 060883 also as COL1152). It was recovered as sister to L. monteithi in a larger clade containing all species within the L. monteithi species group. Other specimens from Alexandra Range and Emmagen Creek have also been sequenced (25- 060882 and 25-057618). Lepanus reidi and L. monteithi are the only species in the group that have metatibiae with a notch on the inner edge and small tufts of setae on the apices of both parameres. Some specimens of L. reidi can be interpreted as having only 2 teeth on the protibiae, but can be separated from L. monteithi by the characters in the key.

Figures

FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe.

FIGURE 9. Habitus of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 1 mm.

FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 0.2 mm.

FIGURE 11. Pygidial configuration of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 1 mm.

FIGURE 12. Map of known records for A, Lepanus monteithi; B, Lepanus reidi including predicted distribution estimated with BIOCLIM models. QLD, Queensland.