Heser nilicola (O. Pickard-Cambridge, 1874)

Senglet, Antoine, 2012, Civizelotes new genus, and other new or little known Zelotinae (Araneae, Gnaphosidae), Revue suisse de Zoologie 119 (4), pp. 501-528 : 521-525

publication ID

https://doi.org/ 10.5962/bhl.part.150203

publication LSID

lsid:zoobank.org:pub:5445CFB2-3480-4E0B-827F-7B15785EE647

persistent identifier

https://treatment.plazi.org/id/03ED87D9-A901-FF95-FF4D-7C84FCF64330

treatment provided by

Carolina

scientific name

Heser nilicola
status

 

Heser nilicola -group

DEFINITION: In these species the apical part of the tegulum is grooved to support the embolus; in the type species the embolus rests on the prolateral part of the median apophysis. Apical grooved sector sometimes partly sclerotized. Long embolus curled left (conter-clockwise) on the left palp. Palpal tibia and retrolateral apophysis short. Embolar base variably reduced. No intercalary sclerite present; the base of the embolar base (Figs 69, 77, 79) can be mistaken for a intercalary. Median apophysis large, except in H. hispanus . The glandular ducts (blind paramedian ducts) are replaced by scattered glands (Figs 75, 83, 89).

SPECIES INCLUDED: Heser nilicola (O. P.- Cambridge, 1874), H. schmitzi ( Kulczynski, 1899) , H. incisupalpis Levy, 1998 , H. bernardi Marinaro, 1967 , H. bonneti Marinaro, 1967 and H. hispanus sp. n.

KEY TO SPECIES OF THE HESER NILICOLA -GROUP Ƌ............................................................ 1 ♀ ............................................................ 3

1a Median apophysis small (Figs 85-86).................. H. hispanus sp. n.

1b Median apophysis large (Figs 70, 77)............................... 2

2a Embolar base reduced; embolus arising from basal prolateral tegular sector (Figs 76, 79).................................... H. schmitzi

2b Embolar base with a projection; embolus arising from dorsal tegular sector (Figs 71-72)..................................... H. nilicola

3a Epigynal plate elongated or triangular (Figs 81-82, 87)................. 4

3b Epigynal plate rounded (Figs 73-74)....................... H. nilicola

4a Epigynal plate triangular, widened almost to the anterior anchoring pockets width (Fig. 87)............................. H. hispanus sp. n.

4b Epigynal plate with more or less parallel sides (Figs 81-82)...... H. schmitzi

Heser nilicola (O. P.- Cambridge, 1874) Figs 69-75 Prosthesima nilicola O. P.- Cambridge, 1874: 380, pl. 5, fig. 8 (description of Ƌ)

For previous synomymy see Platnick (2012).

MATERIAL EXAMINED: GREECE, Peloponnesus, Laconia, south-west of Mavrovouni, litter on sand, 36°43'N 22°32'E; 3 Ƌ, 2 ♀ (last moults of ♀ 09.06. and 02.07.2011); 26.05.2011 GoogleMaps . – Cyclades, Paros, Parikia ; 3 Ƌ, 2 ♀; 25.06.1968 . – Parikia, 37°05'N 25°09'E; 1 Ƌ, 1 ♀ (last moults of Ƌ 29.06, of ♀ 05.06.1998); 01.06.1998. – Naxos, Kato Potamia, 37°06'N 25°26'E; 1 ♀; 10.06.1998. – Crete, Chania, Sougia, 35°15'N 23°48'E; 1 Ƌ (last moult 22.03.2000); 09.10.1999 GoogleMaps . – FRANCE, Corsica, Sud Corse, Testa pass / Bonifacio , 69 m; 1 Ƌ; 27.05.1971 . – Acorane bridge /Sartène; 1 Ƌ; 28.05.1971. – Same; 1 ♀; 19.06.1999. – Portigliolo /Propriano, below vegetation & stones; 2 ♀; 17.06.1999. – ITALIA, Sardinia, Sassari, Palau ( Liscia river ); 1 ♀; 08.09.1968 . – Castelsardo; 1 ♀; 26.05.1999. – Stagno di Calich / Alghero; 1 Ƌ, 1 ♀ (last moults of Ƌ 05.06, of ♀ 01.06.1999); 29.05.1999 . – Nuoro, Cala Ginepro / Orosei, below Juncus ; 1 ♀; 10.06.1999 . – Oristano, Iz Arénas / Narbolia, dry leaf litter on sand; 1 Ƌ, 1 ♀; 31.05.1999. – Stagno San Giusta, Eucalyptus bark; 1 Ƌ, 1 ♀; 01.06.1999. – Cagliari, Quartu (laguna); 1 ♀; 03.06.1999.

DESCRIPTION: See Platnick & Shadab (1983: 274, figs 316, 327-328).

REMARKS: The cleared Ƌ palp shows a relatively well developed embolar base with a projection (Figs 69-72). The base of the embolar base may be mistaken for an intercalary sclerite. ♀: The epigynal plate is rounded. The distance between epigynal plate and anterior anchoring pockets is highly variable. Greek form (Fig. 73): Platnick & Shadab (1983: figs 265-266, showing a faulty connection of the median vulval duct); Chatzaki et al. (2003: fig. 85); FitzPatrick (2007: Fig. 95). Occidental form (Fig. 74, FIGS 69-75

Hesernilicola (O. P.-Cambridge). (69) Leftmalepalp, prolateralview. (70) Id., ventralview. (71) Id., retrolateralview. (72) Id., cleared, dorsalview. (73) Epigynum, fromGreece. (74) Id., from Sardinia. (75) Vulva, dorsal view (Sardinia). Bold lines indicate epigynal folds. Scale 0.2 mm.

Sardinia): Cornick etal. (2004: fig. 3a); Melic (1995: figs 1-2, showingafaulty connection of the median vulval ducts): Sardinia, Corsica, south of France and north Spain.

Heser schmitzi ( Kulczynski, 1899) Figs 76-83 Prosthesima schmitzii Kulczynski, 1899: 359 , pl. 6, fig. 32 (description of ♀).

Zelotes schmitzi . – Platnick & Murphy, 1998: 118, figs 1-4 (♀, description of Ƌ).

MATERIAL EXAMINED: SPAIN, Levant / Murcia, Alicante, Elche , palm grove, 38°17'N 00°42'W GoogleMaps ; 1 Ƌ; 15.05.2002. – Elda , 38°30'N 00°47'W GoogleMaps ; 1 Ƌ; 19.06.1971. – Murcia, Archena , 38°07'N 01°17'W GoogleMaps ; 1 Ƌ; 17.05.2002. – Andalusia, Almeria, Adra ( LaAlbufera ), cultivateddryed pond, 36°46'N 02°58'W GoogleMaps ; 10 Ƌ, 3 ♀ (last moults of 2 Ƌ 27.05 and 15.06, of 2 ♀ 30. 05 and 12.06.2002); 25.05.2002. – Malaga, TorredeMar , 36°44'N 04°07'W GoogleMaps ; 1 ♀; 27.07.1969.

DESCRIPTION: Prosoma and legs brown, with lighter tarsi. Opisthosoma black, with adpressed hairs and black bristles. Leg spination: Femora IV r001; tibiae III r111, IV p110; metatarsi I v000, IV v220/222. Tarsi I, II scopulate.

FIGS 76-83

Heserschmitzi (Kulczynski). (76-80) Leftmalepalp. (76) Prolateralview. (77) Ventralview. (78) Retrolateralview. (79) Cleared, prolateral-ventralview. (80) Id., dorsalview. (81-82) Two epigyna, same population. (83) Vulva, dorsal view. Bold lines indicate epigynal folds. Scale 0.2 mm.

Ƌ: Total length 3.05. Prosoma: 1.36 long, 1.03 wide, 0.54 wide at level of posterior eyes. Eye sizes and interdistances: AME 0.04, ALE 0.07, PME 0.06, PLE 0.06; AME-AME 0.03, AME-ALE 0.01, PME-PME 0.02, PME-PLE 0.02, ALE-PLE 0.03. MOQ length 0.10, front width 0.16, back width 0.18. Clypeus: 0.06 from AME, 0.04 from ALE. Pedipalp (Figs 76-80): Patella dorsally longer than tibia. In cleared bulbus (Figs 79-80) a short embolar radix and a vestigial embolar base visible. Median apophysis large. Scutum occupying 40% of opisthosoma length.

♀: Total length 4.70. Prosoma: 1.53 long, 1.14 wide, 0.60 wide at level of posterior eyes. Eye sizes and interdistances: AME 0.04, ALE 0.07, PME 0.06, PLE 0.06; AME-AME 0.03, AME-ALE 0.01, PME-PME 0.04, PME-PLE 0.03, ALE-PLE. MOQ length 0.14, front width 0.27, back width 0.31. Clypeus: 0.04 from AME, 0.03 from ALE. Epigynum (Figs 81-82), epigynal plate variable, with sides parallel to feebly diverging. Vulva (Fig. 83) with scattered glands on the sides.

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Gnaphosidae

Genus

Heser

Loc

Heser nilicola

Senglet, Antoine 2012
2012
Loc

Zelotes schmitzi

PLATNICK, N. I. & MURPHY, J. A. 1998: 118
1998
Loc

Heser schmitzi (

KULCZYNSKI, W. 1899: 359
1899
Loc

Heser nilicola

CAMBRIDGE, O. P. 1874: 380
1874