Civizelotes caucasius (L. Koch, 1866)

Senglet, Antoine, 2012, Civizelotes new genus, and other new or little known Zelotinae (Araneae, Gnaphosidae), Revue suisse de Zoologie 119 (4), pp. 501-528 : 518-521

Civizelotes caucasius -group

DEFINITION: Posterior eye row procurved. Ƌ pedipalp: Strong elongated embolar radix (Fig. 67) terminating in a bowl. Embolar base containing the sperm duct (Fig. 68), reaching straight to the radix bowl. Relict terminal apophysis with a posterior sclerite without distinct connection. Embolus long, curled right (clockwise) on left palp. Atrium in anterior epigynal sector.

FIGS 58-66

Civizelotes ibericus n. sp. (58-62) Left male palp. (58) Prolateral view. (59) Ventral view. (60) Retrolateral view. (61) Cymbium removed, dorsal view. (62) Id., apical view. (63) Vulva, ventral view. (64) Id., dorsal view. (65) Id., median-lateral view. (66) Epigynum. Bold lines indicate epigynal folds. Scale 0.2 mm.

SPECIESINCLUDED: C. caucasius (L. Koch, 1866) , C. sostitialis ( Levy, 1998) .

REMARKS: The structure of the pedipalp (Figs 65-66), with a long embolar radix and a developed embolar base, excludes these species from the Civizelotes gracilis - group. The study of copulatory mechanism ( Senglet, 2004: 95, figs 26-31) shows that the embolar base and radix are penetrating to mid-length of the vulva; radix not figured, but radix bowl visible at base of embolus ( Senglet, 2004: fig. 31).

Civizelotes caucasius (L. Koch, 1866) comb. n. Figs 67-68 Melanophora caucasia L. Koch, 1866: 144 , pl. 6, fig. 87 (description of ♀).

For previous synonymy see Platnick (2012).

MATERIAL EXAMINED: GREECE, Macedonia, Serres, above Oreini , 1200 m, stones, 41°14'N 23°35'E; 5 Ƌ, 3 ♀; 30.06.2011 . – Thessaloniki, westofAghiosVassilios, 40°41'N 23°05'E; 1 Ƌ (lastmoult 17.06.2008); 31.05.2008. – AghiosVassilios , 40°40'N 23°07'E; 2 ♀ (lastmoultsof ♀ 29.07. and 02.08.1998); 14.06.1998 . – EastofApollonia , belowstones, 40°37'N 23°32'E; 7 Ƌ, 3 ♀ (lastmoultsof ♀ 01.07.2008); 01.06.2008 . – Loutra /Volvilake, 40°39'N 23°25'E; 3 Ƌ (lastmoultsof 2 Ƌ 16 and 22.06.1998); 14.06.1998. – Kastoria, northof Kastoria (pass), 40°35'N 21°18'E; 5 Ƌ, 3 ♀ (lastmoultsof 2 ♀ 21. and 25.06.1998); 20.06.1998 . – Thessalia, Larissa, Gonnos, 39°51'N 22°28'E; 1 Ƌ; 19.05.1968 . – StereaHellas, Phthiotidas, north of Malesina , below stones, olive trees, 38°38'N 23°14'E; 2 ♀ (last moults of ♀ 26. 07. and 09.08.2008); 19.06.2008 . – NearMalesina , 38°37'N 23°13'E; 3 Ƌ, 5 ♀; 21.05.1968 . – Same ; 2 ♀; 15.06.1968 . – Same ; 6 Ƌ, 3 ♀ (with vulva in microvial, last moults of Ƌ 23.05.-25.06, of ♀ 02.-23.06.1998); 21.05.1998 . – MtChlomos , 860 m, stones, 38°36'N 23°10'E; 2 Ƌ, 5 ♀ (last moultsof 4 ♀ 16. 06.-01.07.2011); 14.06.2011 . – Same , 980 m, stones, 38°36'N 23°01'E; 3 Ƌ, 3 ♀ (lastmoultsof ♀ 15. 06.-01.07.2011); 14.06.2011 . – Tràgana , 38°37'N 23°07'E; 1 ♀; 21.05.1968 . – Glyphada / Theologos , 38°39'N 23°14'E; 1 ♀; 04.06.1978 . – Theologos , 38°39'N 23°12'E; 3 Ƌ, 3 ♀ (withvulvainmicrovial, lastmoultsof ♀ 29. 05.-01.06.1998); 20.05.1998 . – Eubea, Theologos, 38°29'N 23°47'E; 1 Ƌ; 11.06.2008. – Dystoslake, 38°22'N 24°08'E; 1 Ƌ; 17.06.1981. – Peloponnesus, Laconia , south-westofMonemvasie, stones, 36°40'N 23°01'E; 1 ♀ (lastmoultof ♀ 25.07.2011); 29.05.2011 . – North-eastofGeraki, 37°02'N 22°43'E; 1 Ƌ, 1 ♀ (lastmoultsofƋ 25.06.1998, of ♀ 01.09.1998); 13.05.1998. – Arcadia , eastofLangadas, 830 m, stones, 37°40'N 22°03'E; 3 Ƌ, 2 ♀ (lastmoultsofƋ 09.-27.06, of ♀ 08 and 27.07.2011); 05.06.2011 . – Argolida, PalaiaEpidauros, Pinuslitter, 37°39'N 23°09'E; 1 Ƌ; 01.10.2004. – Same 1 Ƌ; 05.06.1981. – Same ; 2 Ƌ, 2 ♀ (last moults of 1 Ƌ 17.05, of ♀ 14.05 and 05.06.1998); 14.05.1998 . – AbovePlatani , 700 m, 37°48'N 22°30'E; 4 Ƌ, 1 ♀ (lastmoultsofƋ 20.06.-01.07, of ♀ 15.07.2011); 21.05.2011 . – Corinthos , eastofAngelokastro, stones, 37°45'N 23°01'E; 1 ♀ (lastmoultof ♀ 07.07.2011); 24.05.2011 . – Cyclades, Paros, StaMaria / Naoussa , Salicornia , 37°08'N 25°17'E; 1 ♀; 02.06.1998 . – Naxos , north-eastofSangri, 37°03'N 25°27'E; 1 ♀ (last moultof ♀ 18.06.1998); 05.06.1998 . – NorthofAghiosProkopios, dry, belowstones, 37°05'N 25°21'E; 1 ♀; 06.06.1998. – Koronido , 650-750 m, 37°09'N 25°37'E; 5 ♀ (lastmoults 28. 06.- 03.07.1998); 08.06.1998 . – Crete, Irakleion, Damasta ; 1 ♀; 28.06.1970 . – Chàrakas, Eucaliptus bark, 35°01'N 25°08'E; 1 Ƌ (lastmoult 22.03.2000); 01.10.1999. – Lassithi, Kaminàkion ; 1 Ƌ, 1 ♀; 13.07.1970 . – SPAIN, Nueva Castilla / La Mancha, Albacete, La Gineta ( rio Jucar ), fine leaflitter, 39°10'N 01°58'W; 2 Ƌ (lastmoults 10 and 16.07.2002); 16.05.2002 . – CiudadReal, Fuencaliente , 38°23'N 04°18'W; 3 ♀; 03.08.1969 . – Ruidera , 38°56'N 02°51'W; 1 ♀; 07.08.1969 . – Estremadura, Badajoz, southofVentadelCulebrin / Monesterio, 37°58'N 06°14'W; 1 Ƌ (lastmoult 19.06.1969); 19.06.1969 . – Andalusia, Granada, LaVidriera / Ptodel Pinar , 38°03'N 02°34'W; 1 ♀ (lastmoultof ♀ 05.07.2002); 20.05.2002 . – RoadPuertodela Ragua, pineforest, 37°09'N 03°03'W; 1 Ƌ (lastmoult 10.07.2002); 24.05.2002. – Colladodel Muerto (S. Nevada), 1450 m, 37°08'N 03°28'W; 2 Ƌ (lastmoults 13 and 22.07.2002); 29.05.2002. – Cordoba, AlmodóvardelRio (Breñadam), 37°50'N 05°04'W; 1 Ƌ; 01.06.2009. – Sevilla, Pintadodam / CazalladelaSierra , 37°59'N 05°57'W; 1 Ƌ, 1 ♀; 02.07.1969 . – Huelva, SantaOlalla (rioCala), 37°55'N 06°11'W; 1 Ƌ; 04.07.1969. – PORTUGAL, BeiraAlta, Guarda, Maceira / FornosdeAlgodres (CasaldoMonte), 40°44'N 07°24'W; 1 Ƌ; 09.08.1971 . – FIGS 67-68

Civizelotes caucasius (L. Koch) , left male palp, cleared. (67) Apical view. (68) Ventral-prolateral view.

BULGARIA, Centre , Pazardzik, Pestera; 1 ♀ ; 23.07.1972. – IRAN, Caspian See, Mazanderan, near Dasht , 37°19'N 56°04'E; 1 ♀ ; 27.07.1974.

DESCRIPTION: See Grimm (1985: 281, figs 231, 234-235).