Drassyllus Chamberlin, 1922

Senglet, Antoine, 2012, Civizelotes new genus, and other new or little known Zelotinae (Araneae, Gnaphosidae), Revue suisse de Zoologie 119 (4), pp. 501-528 : 504-507

Genus Drassyllus Chamberlin, 1922

TYPE SPECIES: Drassyllus fallens Chamberlin, 1922 , original designation.

Drassyllus villicoides ( Giltay, 1932) comb. n. Figs 13-19 Zelotes villicoides Giltay, 1932: 28 , fig. 18 (description of Ƌ). – Giltay, 1933: 6, pl. 3, fig. 4 (Ƌ).

MATERIAL EXAMINED: GREECE, Macedonia, Thessaloniki, east of Chortiatis , stones in pasture, 40°37'N 23°07'E; 1 Ƌ (lastmoult 02.06.2005); 07.09.2004 . – WestofAghiosVassilios, 40°40'N 23°05'E; 1 Ƌ, 2 ♀ (lastmoultsofƋ 16.06.2005, of ♀ 08 and 14.05.2005); 08.09.2004. – AghiosVassilios , 40°40'N 23°07'E; 1 ♀; 13.05.1968 . – LoutraApollonias , litter, 40°39'N 23°24'E; 1 Ƌ, 1 ♀ (lastmoultsofƋ 23.05.2005, of ♀ 09.07.2005); 09.09.2004 . – StereaHellas, Phthiotidas , westofMalesina, 38°37'N 23°13'E; 6 Ƌ, 4 ♀ (lastmoultsofƋ 16. 04.-15.05.2005, of ♀ 16. 04.-28.05.2005); 25.09.2004 . – NearMalesina , 38°37'N 23°13'E; 1 ♀; 21.05.1998 . – Peloponnesus, Laconia , south-westofMavrovouni, litteronsand, 36°43'N 22°32'E; 1 ♀; 26.05.2011 . – Argolida, eastofLigourion, Lentiscuslitter , 37°38'N 23°04'E; 1 Ƌ, 3 ♀ (last moultsofƋ 24.04.2005, of ♀ 17. 04.-24.05.2005); 01.10.2004 .

DESCRIPTION: Prosoma dark brown, with tiny hairs. Opisthosoma black, with short adpressed hairs. Legs dark brown except for tawny-yellow tarsi and metatarsi. Tarsi and metatarsi I, II scopulate. Average size about 3/4 that of D. praeficus .

Ƌ: Total length 5.00. Prosoma: 1.95 long, 1.62 wide, 0.77 wide at level of posterioreyes. Eyesizesandinterdistances: AME 0.07, ALE 0.08, PME 0.08, PLE 0.07;

FIGS 11-12

Zelotes balcanicus Deltshev. (11) Left male palp, cymbium removed, anterior-apical view. (12) Id., dorsal view. Scale 0.2 mm.

AME-AME 0.03, AME-ALE 0.01, PME-PME 0.03, PME-PLE 0.04, ALE-PLE 0.06. MOQ length 0.20, front width 0.37, back width 0.43. Clypeus: 0.11 from AME, 0.07 from ALE. Retrolateral tibial apophysis bent dorsally, shorter than dorsal length of tibia. Pedipalp (Figs 13-16): Tibia short; its dorsal length about equal to lateral width. Patella dorsally longer than tibia. Scutum occupying 1/3 of opisthosoma length. Close to D. praeficus , differing by the thin narrow triangular terminal apophysis and the slender embolar radix apophysis (Figs 14, 16); in D. praeficus (Fig. 20) the latter is stronger and more cone-shaped.

♀: Total length 5.66. Prosoma: 2.10 long, 1.58 wide, 0.80 wide at level of posterior eyes. Eye sizes and interdistances: AME 0.07, ALE 0.10, PME 0.08, PLE 0.08; AME-AME 0.08, AME-ALE 0.01, PME-PME 0.02, PME-PLE 0.05, ALE-PLE 0.05. MOQ length 0.24, front width 0.37, back width 0.43. Clypeus: 0.11 from AME, 0.06 from ALE. Epigynum and vulva (Figs 17-19). Anterior anchoring pockets at a distance from the anterior epigynal margins; in D. praeficus (Fig. 21) pocket pressed to the external margin of the epigynum. Modest widening of the lateral fold; median fold reaching the spermatheque in median-anterior position, there covered by cuticle (Figs 18-19); in D. praeficus (Fig. 21) posterior strong widening of the lateral fold; median fold reaching the spermatheque in uncovered lateral-anterior position.

BIOLOGY: Adult males can be collected from April to June.

Drassyllus praeficus (L. Koch, 1866) Figs 20-21 Melanophora praeficus L. Koch, 1866: 155 , pl. 6, figs 97-99 (description of Ƌ and ♀).

For previous synonymy see Platnick (2012). MATERIAL EXAMINED: SWITZERLAND, Vaud, Vich / Nyon ; 1 Ƌ, 1 ♀; 01.-30.05.1996 .

– SPAIN: Lac de Sanabria , 42°06'N 06°41'W and 42°07'N 06°44'W; 1 ♀; 26.08.1971 . – Avila, Road to parking Gredos / Hoyo del Espino , 1470 m, 40°18'N 05°12'W; 1 Ƌ (last moult 28.12.2009); 18.06.2009 . – Andalusia, Granada, Prado de Zangarrilla (S. Nevada), stones, pasture, 2000 m., 37°07'N 03°26'W; 1 ♀; 29.05.2002 . – FRANCE, Provence / Cote d'Azur, Var, Montauroux; 1 ♀; 07.06.1971 . – Languedoc, Pyrénées Orientales, Prats de Mollo , 700 m., 42°24'N 02°25'E; 3 ♀; 29.06.2001 . – Latour de Carol , 42°28'N 01°53'E; 1 ♀; 06.07.2001 . – Above Mas Franco / Enveigt, 42°30'N 01°53'E; 1 ♀; 06.07.2001. – ANDORRA, La Rabassa, 2000 m, 42°26'N 01°32'E; 2 ♀ (last moult of 1 ♀ 24.02.2002); 08.07.2001 . – West of St Julià FIGS 13-19

Drassyllusvillicoides (Giltay). (13-16) Leftmalepalp. (13) Prolateralview. (14) Ventralview. (15) Retrolateralview. (16) Cymbiumdiscarded, retrolateralview. (17) Epigynum. (18-19) Two vulvae, dorsal view, same population.

FIGS 20-21

D. praeficus . (20) Left male palp, cymbium removed, retrolateral view. (21) Left side of vulva, ventral view. Bold lines indicate epigynal folds. Scale 0.2 mm.

deLoria, schistlitter, 42°27'N 01°28'E; 3 ♀ (lastmoultsof 2 ♀ 05 and 08.06.2004); 27.05.2004. – GREECE, Macedonia, Serres , aboveOreini, 1200 m, stones, 41°14'N 23°35'E; 1 ♀; 30.06.2011 . – Thessaloniki , westofAghiosVassilios, 40°40'N 23°05'E; 1 Ƌ, 1 ♀ (lastmoultsof Ƌ 11.06, of ♀ 09.06.2005); 08.09.2004 . – Same ; 4 ♀; 31.05.2008 . – EastofAghiosVassilios, litter, 40°40'N 23°07'E; 2 Ƌ (lastmoults 17 and 20.06.2005); 08.09.2004. – Same ; 2 Ƌ, 4 ♀; 13.05.1968 . – Same ; 1 ♀; 28.05.1982 . – Same , 40°40'N 23°07'E; 4 ♀; 14.06.1998 . – Thessaloniki , 40°43'N 22°58'E; 1 ♀; 13.-18.05.1968 . – Kastoria , northofKastoria (pass), 40°35'N 21°18'E; 1 ♀; 20.06.1998 . – Peloponnesus, Achaia, KatoAlissos , 38°09'N 21°35'E; 3 ♀ (lastmoultof 1 ♀ 10.05.1998); 29.04.1998 . – Arla , 38°04'N 21°36'E; 1 ♀ (lastmoultof ♀ 05.05.1998); 30.04.1998 . – Argolida, eastofMyli / Nauplie , leaflitter, 37°35'N 22°44'E; 1 ♀ (lastmoult 03.06.2005); 01.10.2004 . – Cyclades, Paros, StaMaria / Naoussa , ( Salicornia ), 37°08'N 25°17'E; 1 ♀; 02.06.1998 . – Crete, Chania, Omaloshautplateau , stones, 35°20'N 23°53'E; 2 Ƌ, 2 ♀ (lastmoultsofƋ 01.01 and 04.01, of ♀ 31.12.1999); 07.10.1999 . – Episkopi , litter, soilcrevices, 35°30'N 23°46'E; 1 ♀ (lastmoult 14.01.2000); 10.10.1999 . – Rethimnon, Koxare / Spili , irigatedvegetation, 35°14'N 24°28'E; 1 Ƌ, 2 ♀ (lastmoultsofƋ 30.12.1999, of ♀ 08.01.2000); 03.10.1999 . – Choumerion / Pérama , 35°21'N 24°44'E; 1 Ƌ (lastmoult 04.01.2000); 05.10.1999 . – Missiria / Rethimnon , leaflitter, soilcrevices, 35°22'N 24°31'E; 2 Ƌ; 06.10.1999 . – Lassithi , KatoMetochiLassithiou, 35°11'N 25°26'E; 2 Ƌ, 2 ♀ (withpalpusin microvial, lastmoultsofƋ 25 and 30.12.1999, of ♀ 01. 01 and 06.01.2000); 25.09.1999 . – ROMANIA, Dobrogea, Constanca, Baneasa ; 1 Ƌ (withpalpusinmicrovial); 30.05.1972 . – Adamclisi ; 1 Ƌ; 31.05.1972 . – Mangalia ; 2 ♀; 02.06.1972 . – Techirghiol ; 1 ♀; 03.06.1972 . – BULGARIA, Razgrad, Hlebarovo ; 1 ♀; 08.07.1972 . – Varna, SlatniPjasaci ; 1 ♀; 11.07.1972 . – Burgas, Obzor ; 3 ♀; 14.07.1972 . – Karnobat ; 2 ♀; 15.07.1972 . – Plovdiv, Backovo ; 1 ♀; 21.07.1972 . – IRAN, Esfahan, Lorestan, Dizgarán , 33°44'N 46°59'E; 3 ♀; 16.05.1974 . – Fars, nearGhaderábád, 30°22'N 53°18'E; 1 ♀; 11.06.1974 . – Khorasan, eastofChamanBid, 37°26'N 56°37'E; 1 ♀; 14.07.1974 .

DESCRIPTION: SeeGrimm (1985: 267, figs. 317, 319-320).