Monacha (Metatheba) samsunensis, , Clauss, 1990

Hausdorf, Bernhard, 2000, genus Monacha in the Western Caucasus Gastropoda: Hygromiidae), Journal of Natural History 34, pp. 1575-1594 : 1591-1592

Monacha (Metatheba) samsunensis (L. PfeiOEer, 1868)

(®gures 7, 8)

Helix samsunensis L. PfeiOEer, 1868: 480. Locus typicus:`Samsun’, Turkey.

Helix (Carthusiana) frequens, Retowski 1889: 237 [partim, non Mousson, 1859].

Theba orientalis Hesse, 1914: 261 [partim]. Locus typicus (restrictus, Baker 1963):`Tokat’, Turkey.

Theba scrobiculosa Lindholm, 1922: 360 . Locus typicus:`Kordsul (Gouv. Batum) auf der landwirtschaftlichen Station ``Kardsweli ’’ ’.

Theba scrobiculosa var. perforata Lindholm, 1922: 360 . Locus typicus:`Kordsul (Gouv. Batum) auf der landwirtschaftlichen Station ``Kardsweli ’’ ’.

Theba (Metatheba) orientalis subcarthusiana var. scrobiculosa, Likharev and Rammelmeier 1952: 434 .

Monacha (Metatheba) subcarthusiana scrobiculosa, Schileyko 1978: 291 .

Shell (®gure 7, 8). Depressed conical±globular; with 5±6 convex whorls; teleoconch with wrinkled growth-ridges; upper whorls usually, rarely also lower whorls, with hair-scars (or very short hairs); especially lower whorls with Ô distinct incised spiral striae; whitish or light corneous, then a whitish peripheral band might be discerned; body whorl rounded; aperture elliptical; upper insertion of the peristome usually distinctly descending; peristome sharp, hardly expanded, with a whitish internal rib; umbilicus narrow or very narrow, partly or almost completely obscured by the reēcted columellar edge.

Measurements. Various localities (n 5 30): D: 11.0± 19.1 mm, x Å 5 14.4 mm; H: 6.7±13.0 mm, x Å 5 9.7 mm; D/H: 1.37±1.65, x Å 5 1.49.

Genitalia. The penial papilla is conical or cylindrical. The two little rami®ed glandulae mucosae insert near the proximal end of the vagina. The bursa of the bursa copulatrix is Ô rounded triangular or oval. The right ommatophora l retractor passes to the left of the genitalia. The variation of the proportions of some sections of the genitalia is compiled in table 4.

Remarks. The types of T. scrobiculosa Lindholm correspond with M. samsunensis . Lindholm (1922) compared his species with `Th. samsunensis ’ from Gagra. However, the specimens from Gagra do not belong to M. samsunensis but to M. perfrequens . As Lindholm (1922) already recognized, the types of T. scrobiculosa Lindholm (the true M. samsunensis ) diOEer from M. perfrequens in the wider umbilicus. The identity of T. scrobiculosa with M. samsunensis should be checked anatomically; T. scrobiculosa is certainly not closely related to M. subcarthusiana as supposed by Likharev and Rammelmeier (1952) and Schileyko (1978). It diOEers from M. subcarthusian a in the corneous shell with distinct hair-scars.

The type locality of T. scrobiculosa in the former district of Batum could not be localized. Some parts of the former district of Batum belong to the Turkish Vilayet Artvin today. Because there are no other known localities of M. samsunensis in Georgia, it is not clear whether this species occurs in Georgia. M. samsunensis is known from the Vilayet Artvin (Hausdorf, in press).

The specimens from the debris of the Chorokh from Batumi (SMF 69925, 69926) recorded by Retowski (1889) probably also belong to M. samsunensis .

Distribution and material. Monacha samsunensis is distributed in northern and central Turkey from the Vilayet Bilecik in the west to the Vilayet Artvin in the east. Its occurrence in Adzhariya is questionable.

Kordsul at the agricultural station Kardsveli (lectotype of scrobiculosa (design. Schileyko 1978): ZIRAN; lectotype of scrobiculosa var. perforata (design. nov.):

ZIRAN, measurements: D5 11.4 mm, H5 7.4 mm; paralectotypes of scrobiculosa var. perforata: ZIRAN ).