Monacha (Metatheba) ciscaucasica, Hausdorf, 2000

Hausdorf, Bernhard, 2000, genus Monacha in the Western Caucasus Gastropoda: Hygromiidae), Journal of Natural History 34, pp. 1575-1594 : 1586-1589

Monacha (Metatheba) ciscaucasica , sp. n.

(®gures 5, 14, 19)

Helix carthusiana [sic], Kaleniczenko 1853: 83 [partim, non O. F. MuÈller, 1774].

Helix frequens, Mousson 1863: 318 [partim, non Mousson, 1859].

Helix frequens, Rosen 1911: 101 [partim, non Mousson, 1859].

Theba orientalis roseni Hesse, 1914: 263 [partim, non Hesse, 1914].

Shell (®gure 5). Depressed conical±globular; with 4×±5× convex whorls; teleoconch with wrinkled growth-ridges and Ô distinct incised spiral striae; whitish or light corneous, then a whitish peripheral band might be discerned; body whorl rounded; aperture elliptical or almost circular; upper insertion of the peristome usually slightly descending; peristome sharp, not expanded, with a whitish internal rib; umbilicus narrow, slightly obscured by the reēcted columellar edge.

Measurements. Pyatigorsk, Mashuk Gora (n 5 20): D: 8.3±13.9 mm, x Å 5 11.4 mm; H: 5.7±9.4 mm, x Å 5 7.6 mm; D/H: 1.43±1.58, x Å 5 1.50.

Genitalia (®gure 14). The penial papilla is Ô conical or almost cylindrical. The two glandulae mucosae are simple or little rami®ed. The bursa of the bursa copulatrix is Ô oval. The right ommatophoral retractor crosses with the penial retractor. The measurements of some sections of the genitalia are compiled in table 3 and the variation of the proportions of some sections of the genitalia is compiled in table 4.

Remarks. Monacha ciscaucasica cannot be distinguished from M. (Paratheba) roseni by shell characters. The umbilicus of M. ciscaucasica is narrower than the umbilicus of M. subcarthusiana , but wider than the umbilicus of M. perfrequens .

The genitalia of M. ciscaucasica diOEer from the other Caucasian Metatheba species , e.g., in the length of the penial papilla which is in relation to the epiphallus longer than in M. subcarthusiana , but shorter than in M. perfrequens .

Etymology. Because of the occurrence in Ciscaucasia.

Type locality. Russia: Pyatigorsk , Mashuk Gora, LP47 .

Distribution and material (®gure 19). So far M. ciscaucasica is only known from the surroundings of Pyatigorsk in the Northern Caucasus. Only the specimens from the Mashuk Gora are paratypes.

Russia. Pyatigorsk , Mashuk Gora, LP47 (det. anat.; holotype ZMH 2771 , leg. E. Clauss, measurements: D 5 12.9 mm, H 5 8.8 mm; CLA; SMF 278702 ; ZMH 2770 ); Pyatigorsk, LP47 (paralectotype s of roseni: SMF 6605; ZMZ 505714 (partim?) specimens to Mousson 1863; SMF 69919, 69927 , 98770 ; ZIRAN; ZMUM 4519 , 8424 ) .

( Metatheba ) species. Abbreviations: ep5 epiphallus; ¯ 5 ¯agellum; p 5 penis; pp5

penial papilla; v 5 vagina up to the glandulae mucosae; v 5 total length of the

gm t

vagina.