Cephennodes (Cephennodes) yangrianus, Jałoszyński, 2017

Jałoszyński, Paweł, 2017, Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4349 (1), pp. 1-120 : 26-27

Cephennodes (Cephennodes) yangrianus sp. n.

( Figs 36 , 52, 72–73, 89)

Type material. Holotype: NEPAL (Sindhupalchowk District): ♂, one label: " NEPAL (Prov. Bagmati) / Yangri Ridge / 4350 m, 22.IV.81 / Löbl & Smetana" [white, printed] ( MHNG) . Paratypes (15 exx): 4 ♀♀, same data as for holotype ; 1 ♂, " NEPAL (Prov. Bagmati) / Yangri Ridge / 4150 m, 24.IV.81 / Löbl & Smetana" [white, printed] ; 2 ♂♂, " NEPAL (Prov. Bagmati) / Yangri Ridge / 4200 m, 21.IV.81 / Löbl & Smetana" [white, printed] ; 1 ♀, " NEPAL (Prov. Bagmati) / Yangri Ridge / 4300 m, 23.IV.81 / Löbl & Smetana" [white, printed] ; 7 ♂♂, " NEPAL (Prov. Bagmati) / Yangri Ridge / 4700–4800 m, 22.IV.81 / Löbl & Smetana" [white, printed] (paratypes in MHNG and cPJ).

Diagnosis. BL 1.23–1.31 mm; elytra long, EI 1.13–1.22; body brown with much darker, dark chocolate head and pronotum; antennae unmodified; protibiae in male not thickened but slightly curved inwards, with long patch of dense spatulate setae; apical projection of median lobe in ventral view slightly broadening distally, in lateral view very weakly recurved; each paramere with two strikingly broad and long apical setae which are nearly equal in length.

Description. Body of male ( Fig. 36 ) dark brown with much darker, dark chocolate head, pronotum and antennae, also elytral suture slightly darkened; relatively slender, with shallow constriction between pronotum and elytra, moderately convex, covered with light brown setae; BL 1.23–1.30 mm (mean 1.26 mm).

Head subtrapezoidal, HL 0.18–0.20 mm (mean 0.18 mm), HW 0.29–0.33 mm (mean 0.30 mm); vertex and frons confluent and weakly convex; supraantennal tubercles distinct but weakly elevated; eyes moderately large, strongly convex, coarsely faceted. Punctures on frons and vertex small, superficial and diffuse, inconspicuous; setae very short, sparse, suberect. Antennae moderately long and moderately thickened, AnL 0.60–0.63 mm (mean 0.61 mm); antennomeres moderately compactly assembled, pentamerous club indistinctly delimited and about as long as half AnL, surface of five terminal antennomeres less glossy than that of proximal antennomeres; antennomere I about 1.3 × as long as broad; II distinctly narrower but only slightly shorter than I, 1.6 × as long as broad; III and IV subequal in shape, each slightly narrower than II and much shorter, about as long as broad, V indistinctly broader and longer than IV, about as long as broad, VI indistinctly broader than V and slightly shorter, slightly transverse, VII distinctly broader and slightly longer than VI, distinctly transverse, VIII slightly broader but slightly shorter than VIII, strongly transverse, IX and X each gradually larger than preceding one and distinctly transverse, XI distinctly broader than X, about as long as IX–X together, about 1.7 × as long as broad.

Pronotum in dorsal view subrectangular, broadest near middle, PL 0.38–0.40 mm (mean 0.38 mm), PW 0.49– 0.53 mm (mean 0.50 mm); pronotum strongly convex at middle and strongly flattened near each hind corner but the flattened area very small; anterior margin in strictly dorsal view slightly convex; lateral margins not microserrate, weakly rounded on entire length, behind broadest site distinctly convergent toward slightly obtuseangled posterior corners; posterior margin shallowly bisinuate; lateral carinae very narrow, fused with lateral margins; antebasal pits small and shallow, diffuse, each nearly equally distant from posterior and lateral pronotal margin. Punctures on disc fine, inconspicuous, surface glossy; setae short and moderately dense, suberect.

Elytra elongate, oval, broadest between middle and anterior third, EL 0.68–0.75 mm (mean 0.70 mm), EW 0.58–0.63 mm (mean 0.60 mm), EI 1.13–1.22. Humeral calli weakly elevated, elongate; subhumeral lines very short, equal to only about 0.15 EL but relatively distinct, developed as moderately sharp stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron large and located in very short, deep, round basal impression; apices of elytra separately rounded. Punctures more distinct than those on pronotum and dense, but still only superficial and inconspicuous; setae similar to those on pronotum.

Hind wings reduced, shorter than elytra.

Metaventrite with indistinct postmesocoxal impressions, median area covered with fine punctures.

Protibiae (Fig. 52) not thickened but slightly curved inwards, with large mesal patch of dense spatulate setae.

Abdomen and hind legs unmodified.

Aedeagus ( Figs 72–73 ) in ventral view with slightly asymmetrical median lobe broadest near base, AeL 0.38 mm, median lobe in ventral view with nearly straight and nearly symmetrical distal margin; apical projection in ventral view slightly broadened in subapical region, in lateral view weakly recurved, so that apex is directed distodorsally; each paramere with two strikingly massive, thickened and very long setae.

Female. Externally similar to male, protibiae nearly straight, but also with mesal patch of dense setae; BL 1.25–1.31 mm (mean 1.30 mm); HL 0.18 mm, HW 0.30–0.31 mm (mean 0.30 mm), AnL 0.56–0.58 mm (mean 0.57 mm); PL 0.38–0.43 mm (mean 0.39 mm), PW 0.51–0.55 mm (mean 0.53 mm); EL 0.70–0.75 mm (mean 0.73 mm), EW 0.63–0.65 mm (mean 0.63 mm), EI 1.12–1.20.

Distribution ( Fig. 89 ). Central Nepal.

Etymology. Locotypical, after Yangri Ridge in Nepal.

Remarks. This species is most similar to C. meredaranus , both are small and have the elytra lighter than the pronotum, head and antennae; and males have the protibiae slender, not thickened. The protibiae in C. yangrianus , however, are slightly curved and bear a large patch of spatulate setae, whereas those of C. meredaranus are covered with golden setae without the mesal setal patch. Aedeagal structures also differ between these two similar species.

Figures

FIGURES 32 – 39. Dorsal habitus of Cephennodes (s. str.) bagmatianus sp. n. (32), C. (s. str.) popeye sp. n. (33), C. (s. str.) clavodentatus sp. n. (34), C. (s. str.) meredaranus sp. n. (35), C. (s. str.) yangrianus sp. n. (36), C. (s. str.) suturalis sp. n. (37), C. (s. str.) karnaliensis sp. n. (38), and C. (s. str.) churtanus sp. n. (39).

FIGURES 72 – 79. Aedeagus in ventral (72, 74, 76, 78) and lateral (73, 75, 77, 79) views. Cephennodes (s. str.) yangrianus sp. n. (72 – 73), C. (s. str.) suturalis sp. n. (74 – 75), C. (s. str.) karnaliensis sp. n. (76 – 77), C. (s. str.) churtanus sp. n. (78 – 79). Abbreviations: ap, apical projection.

FIGURES 88 – 89. Distribution of Cephennodes (s. str.) bagmatianus sp. n. (88, circle), C. (s. str.) popeye sp. n. (88, triangle), C. (s. str.) clavodentatus sp. n. (89, circle), C. (s. str.) meredaranus sp. n. (89, circle), C. (s. str.) yangrianus sp. n. (89, triangle).