Claustropyga caetrata, Vilkamaa, 2024

Vilkamaa, Pekka, 2024, Description of two new species of Claustropyga Hippa, Vilkamaa & Mohrig (Diptera, Sciaridae) from the northern Palaearctic, Zootaxa 5463 (1), pp. 137-144 : 141-143

Claustropyga caetrata sp. n.

Figs 4 B , 5–6

Material studied. Holotype male. RUSSIA, Yamalo-Nenets Autonomous Okrug, on the river Longotegan , 67º32’ N, 66º72’ E, Malaise trap, 2–28.VII.2015, A. Barkalov ( MZH) . Paratype. Same data as holotype, 1 male ( MZH) .

Male. Head. Face and antenna concolorous brown, maxillary palpus pale brown. Eye bridge 2 facets wide. Face with 2–5 scattered strong setae. Clypeus non-setose. Maxillary palpus with 2 segments; 1 st segment longer than 2 nd segment; 1 st segment with 2 sharp setae, with a large indistinct dorsal patch of sensilla. Body of 4 th antennal flagellomere 1.45–1.50x as long as wide, the neck slightly shorter than wide, the longest setae as long as the width of flagellomere. Thorax. Brown, setae dark. Scutum with some long and strong laterals and long dorsocentrals. Scutellum with 4 long and strong setae and with some short setae. Anterior pronotum with 2 setae. Proepisternum with 5 setae. Wing. Length 1.7 mm. Width/length 0.45. Anal lobe small. R 1 /R 0.70. c/w 0.65–0.85. r-m and bM equally long or r-m slightly longer. Fork of M longer than stM. r-m and bM non-setose. Halter pale brown. Legs. Fore coxa yellow, mid- and hind coxae pale brown, coxal setae dark. Fore tibial organ with indistinct patch of few strong setae. Fore tibial spur slightly longer than the tibial width. Abdomen. Pale brown, setae dark, long and strong. Hypopygium ( Fig. 5 ). Pale brown, as abdomen. Tergite IX long. Intergonocoxal area lobe-like produced, with shallow notch; gonocoxa nearly as long as gonostylus, wide, with short medial setosity, with membraneous area non-setose; with short apicoventral elongated seta. Gonostylus ( Figs 4 B , 5 C ) wide, shortly setose, with prominent ventromedial lobe with long and sharp setae; strongly excavated; with straight dorsomedial margin; with short apical tooth with some sharp cusps ( Fig. 6 A ); with 1 megaseta at medial margin of gonostylar lobe (holotype with 2 in one gonostylus), 1 megaseta in medial excavation arising from large basal body. Tegmen ( Figs 5 A , 6 B ). wide basally with distinct basolateral shoulders, strongly narrowed at middle; slightly sclerotized laterally and apically, with a small area of minute aedeagal teeth. Aedeagal apodeme moderate.

DNA. Unknown.

Female. Unknown.

Discussion. The holotype and paratype of Claustropyga caetrata sp. n. differ slightly in the exact form of the tegmen and of the intergonocoxal lobe ( Figs 5 A , 6 B ) but the specimens are considered conspecific. Claustropyga caetrata resembles C. corticis ( Mohrig & Antonowa, 1978) in having a large medial lobe in the gonostylus. It differs in having only two or three gonostylar megasetae instead of five and in having distinct lateral shoulders in the tegmen instead of smoothly curved sides. Claustropyga caetrata differs from all other known species of the genus in having sharp teeth-like cusps in the apical tooth of the gonostylus.

Etymology. The name is a Latin adjective, caetrata , armed with a small shield, referring to the wide ventral lobe of the male gonostylus.

Figures

FIGURE 4. Gonostylus, ventral. A. Claustropyga alveata sp. n. (holotype). B. C. caetrata (holotype). Scale 0.1 mm.

FIGURE 5. Claustropyga caetrata sp. n. (holotype). A. Hypopygium, ventral. B. Hypopygium, dorsal. C. Gonostylus, dorsal. Scale 0.1 mm.

FIGURE 6. Claustropyga caetrata sp. n. (paratype). A. Apical part of gonostylus, ventral. B. Tegmen, ventral. Scale 0.05 mm.