Tillicera soror Schenkling, 1902

Gerstmeier, Roland & Bernhard, Nadine, 2010, Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae), Zootaxa 2359 (1), pp. 1-34 : 29-31

Tillicera soror Schenkling, 1902

( Figs. 16 , 41–42 )

Tillicera soror Schenkling 1902: 127 ; Schenkling 1903: plate 1, fig. 10; Pic 1934: 133 ( Rhytidoclerus ); Corporaal 1939: 26.

Specimens examined: Holotype: Type, Tillicera soror Schklg., British Bootang, L. Durel, Rene Oberthur (RMNH) .

Paratypes: ♀. Tillicera soror, Schenkling, 1902 , ZMAN type COLE.1800.1, Type: Tillicera soror Schklg., Cotype, British Bootang, Maria Basti, L. Durel , ex Mus., Lugd. Bat. (1 ex., ZMAN); ♀. Tillicera soror, Schenkling, 1902 , ZMAN type COLE.1800.2, Type: Tillicera soror Schklg. , coll. R. Oberthür, British Bootang, Maria Basti, 1899, ex Mus., Lugd. Bat. (1 ex., ZMAN).

Others: British Bootang, Padong Himalaya , 1914 rec.: L. Durel, ex coll. R. Oberthur (2 ex., RGCM) ; Inde, Ex Museo, Mniszech , Coll. M. Pic (1 ex., MNHN) ; Assam, India, Ex Museo, E. Hintz, ex Coll. R. Oberthur (1 ex., MNHN) ; British Bootang, Maria Basti , 1900, ex Coll. R. Oberthur (2 ex., MNHN) ; British Bootang , Maria Basti, L. Durel, ex Coll. R. Oberthur (8 ex., MNHN) ; British Bootang, Maria Basti , 1899, ex Coll. R. Oberthur (18 ex., MNHN) ; British Bootang , Maria Basti, L. Durel, Coll. M. Pic (1 ex., MNHN) ; Maria Basti, British Bootang , ex Coll. R. Oberthur (3 ex., MNHN) ; British Bootan, Padong, L. Durel , 1914, ex Coll. R. Oberthur (6 ex., MNHN) ; British Bootan, Padong, L. Durel 1914, Coll. A. Kuwert, Coll. R. Oberthur (3 ex., MNHN) ; Pedong, A. Desgodins , ex Coll. R. Oberthur (1 ex., MNHN) ; Pedong, A. Desgodins , Coll. M. Pic (1 ex., MNHN) ; British Bootang, L. Durel , 1898, ex Coll. R. Oberthur (20 ex., MNHN) ; British Bootang, Sakiou, L. Durel , ex Coll. R. Oberthur (2 ex., MNHN) ; Bhoutan, Anglais, R. Oberthur 1900 (1 ex., MNHN) ; Rongli , 26.– 27.7.1900, ex Coll. R. Oberthur (4 ex., MNHN) ; Khasis, 7.1894, Nat. Coll , ex Coll. R. Oberthur (1 ex., MNHN) ; Khasis, 7.1894, Nat. Coll , Coll. M. Pic (1 ex., MNHN) ; Gnatong , 1.– 2.7.1900, ex Coll. R. Oberthur (1 ex., MNHN) ; Ex Museo, Parry , ex Coll. R. Oberthur (3 ex., MNHN) ; Nepal: Kosi - #13, Depitar 27°27’N, 87°17’E to Barabishe, 27°26’N, 87°18’E, 1250– 560m, 12.6.01, NHMB Basel, expedition to Nepal, 2001 (2 ex., BMNH) ; Nepal: Kosi - #9a, Chichilla, 1900–2000m, 27°28’N, 87°14’E, 3.– 5.6.2001, NHMB Basel, expedition to Nepal, May / June 2001 (1 ex., NHMB) ; India, W Bengalen, 1000 m, Darjeeling Distr. , Kurseong, Malloutar Umg. 6.–14.7., leg. N. Dangal 1989 (1 ex., NMEG) ; Nepal 469 Myagdi Distr. , Myagdi Khola S Boghara, 1400 m, 27.– 28.5.1995, Martens & Schawaller (1 ex., SMNS) ; 413 Sankhua Sabha Distr., Arun Valley betw. Hurure and Chichila , tree, rich cultural land, 2000 m, 17.6.1988 Martens & Schawaller leg., Nepal-Expedition, Jochen Martens (1 ex., SMNS) ; British Bootang, Maria Basti , 1899, ex Mus. R. Oberthur 1922 (1 ex., ZMAN) ; British Bootang , Maria Basti, L. Durel 1899, ex Mus. R. Oberthur 1922 (2 ex., ZMAN) ; British Bootang , Maria Basti, L. Durel, ex Mus. R. Oberthur 1922 (1 ex., ZMAN) ; Gopaldhara, Darjeeling , 3440–4720 ft., H. Stevens, from Foliage, 1925 from, Brit. Mus. (1 ex., ZMAN) ; Gopaldhara , Nepal-Sikhim Frontier, H. Stevens 16.9.18, 1925 from, Brit. Mus. (1 ex., ZMAN) ; Gopaldhara , Nepal-Sikhim Frontier, H. Stevens 1920, 1925 from Brit. Mus. (1 ex., ZMAN) ; Gopaldhara, Bw., Darjiling , 4720 ft. -19., H. Stevens, 1925 from Brit. Mus. (1 ex., ZMAN) ; Ex Chinde, Deprepani 6000, Darjeeling, Bengal, J.C.M. Gardner , 19.4.1930, R.R.D. 22, B.C. R. 483, Cage 192 (1 ex., ZMAN) ; Zool. Surv. Ind., Sureil, 5000 ft., Darjiling dist. E. Himalayas, 11.–31.10.17, N.A. & F.G., 1937, Retained from Indian Museum Calcutta (1 ex., ZMAN) ; Zool. Surv. Ind., Above Tura, Garo hills, Assam , 3500–3900 ft., 15.7.–30.8.17, S. Kemp., on fallen tree, 1937, Retained from Indian Museum Calcutta (1 ex., ZMAN) ; Tonkin, Tugun, Quang , 5.1915, R. Vitalis de Salvaza leg., vend. 1929 (1 ex., ZMAN) .

Length: 9–13mm (101 specimens)

Collecting period: April – October, December

Head: Black, anterior part of clypeus, labrum and palpi brown, posterior part of clypeus black; distributed with fine punctation; cranium vested with dense, predominantly pale, setae; head including eyes slightly broader than anterior width of pronotum.

Antennae: Dark-brown to black, A1 to A4 sometimes slightly paler, with depressed pale and dark setae; antennomeres triangularly dilated from A6 or A7 onwards, A10 broader than long, or as long as wide.

Pronotum: Red-brown to brown, surface anterior to transverse depression mostly black; surface anterior to transverse depression finely and diffusely punctate, posterior of transverse depression more coarsely and irregularly punctate; vested with anteriorly directed and erect dark and pale setae; length:width ratio 1.14:1 (broadest at middle).

Scutellum: Brown, with pale setae, finely punctate.

Elytra: Black; basal third with a red-brown macula (side margins bordered black), margined posteriorly by two yellowish pigmented transverse maculae (spanning from the lateral margin to mid-way towards suture) with dense, pale, depressed setae, between these maculae with a semicircular sickle-like pattern of pale setae, apical third with a pale yellowish pigmented transverse fascia (not always pigmented towards suture), with dense, pale, depressed setae; basal third regularly and deeply punctate with 6 to 7 deep rows, only sometimes extending towards posterior transverse fascia, the black part with fine, dense, punctation, diameter of punctures larger than interstices, apex densely and finely punctate; basal third with longer, erect, pale and dark setae, middle part predominantly with dark, posteriorly directed and semi-decumbent, setae, apex with dark setae gradually more erect approaching apices; length:width ratio 2.24:1.

Legs: Brown to black, base sometimes paler, with dense, erect, pale setae; tibiae with longitudinal carina.

Ventral surface: Abdomen black to brown, thorax brown, with pale, depressed setae; 5 th ventrite of males deeply emarginated.

Aedeagus: Asymmetrical, tooth-like structures distinct, large parameres.

Distribution: Nepal, Bhutan, India.

Differential diagnosis to T. auratofasciata and T. javana : The antennae of T. soror are triangularly (less stronger) dilated from A6 onwards; besides differences in punctation, also the coloration is different; the brown basal macula of T. soror is smaller, and less precisely bordered than that of T. auratofasciata and T. javana ; parameres and pygidium also differ.

Figures

FIGURE 41: Tillicera soror, aedeagus ventral aspect, aedeagus lateral aspect, pygidium and ventrite VI.

FIGURES 13–17: Dorsal habitus of Tillicera spp.. Fig. 13. Tillicera obscura n. sp.. Fig. 14. Tillicera paula. Fig. 15. Tillicera pseudocleroides n. sp.. Fig. 16. Tillicera soror. Fig. 17. Tillicera tonkinensis n. sp..