Lepthercus, Purcell, 1902

Ríos-Tamayo, Duniesky & Lyle, Robin, 2020, The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae) phylogeny and taxonomy, Zootaxa 4766 (2), pp. 261-305: 267-269

publication ID

http://doi.org/ 10.11646/zootaxa.4766.2.2

publication LSID

lsid:zoobank.org:pub:urn:lsid:zoobank.org:pub:BA570D47-FE77-446C-B000-9208C9A6E957

DOI

http://doi.org/10.5281/zenodo.3803502

persistent identifier

http://treatment.plazi.org/id/03F11B20-FFD8-FF80-8EE8-394BFE6FD7F1

treatment provided by

Carolina

scientific name

Lepthercus
status

 

Genus Lepthercus  

http://zoobank.org/ urn:lsid:zoobank.org:act:E3F3DEC2-513B-4D9C-9F4C-06A47508996B

Lepthercus Purcell, 1902: 379   . Type species: Lepthercus dregei   by original designation. Hewitt, 1917: 699. Raven, 1985a: 86.

Diagnosis. Males can be distinguished from the genera Hermacha   and Entypesa   , by possessing a megaspine on a raised cuticular spur on tibia I (absent in Hermacha   and Entypesa   ), thin prolateral spiniform setae on male palp tibia ( Figs. 3E View FIGURE , 5E View FIGURE , 9E View FIGURE , 12E View FIGURE , 14E View FIGURE , 21E View FIGURE ; thick in Hermacha   and Entypesa   ) and presence of a protuberance on metatarsus I of males ( Figs. 5J View FIGURE , 9J View FIGURE , 12J View FIGURE , 14J View FIGURE , 17I View FIGURE , 19J View FIGURE , 21I View FIGURE , 23I View FIGURE ; absent in Hermacha   and Entypesa   ). Lepthercus   differs too by Hermacha   , by the presence of preening combs in both sexes ( Fig. 2A View FIGURE ), and from Entypesa   by the presence of a dense scopula on tarsi I-II (less dense than Hermacha   ).

Description. Fovea short, more or less straight. Clypeus narrow with some bristles on the edge. Eye tubercle raised, well defined. Usually no cuspules on labium (1–3 if present). Maxillae rectangular; anterior lobe rounded, prolateral face of females slightly curved or curved, with a short or long soft area. Serrula present in females ( Fig. 2 View FIGURE D-K); in L. engelbrechti   sp. nov., L. haddadi   sp. nov., and L. sofiae   sp. nov., is present in both sexes. Maxillae with numerous cuspules on inner corner, not on mound (minute in males of L. kwazuluensis   sp. nov., L. confusus   sp. nov. and L. mandelai   sp. nov.; Fig. 2C View FIGURE ). Postlabial sigilla consisting of a pair of well-marked depressions, almost meeting in the center. Sternum covered with black hairs and bristles, with posterior sigilla marginal, small and well defined. Chelicerae without rastellum. Males with a small and setose intercheliceral tumescence ( Figs. 3C View FIGURE , 5C View FIGURE , 9C View FIGURE , 12C View FIGURE , 14C View FIGURE , 17C View FIGURE , 19D View FIGURE , 21C View FIGURE , 23C View FIGURE ; absent in females).

Leg formula 4123; all legs covered with sparse hairs. Cymbium short without spines; palp tarsi of females with two basal spines. Palp tibia of males normal (Group dregei, Fig. 3D View FIGURE ) or wide, dorsally very convex (Group haddadi, Fig. 17D View FIGURE ); slender in Hermacha   and Entypesa   . Thin spiniform setae usually present in the prolateral side of the palp tibia of males (Figs. Figs. 3E View FIGURE , 5E View FIGURE , 9E View FIGURE , 12E View FIGURE , 14E View FIGURE , 21E View FIGURE ; absents in L. engelbrechti   sp. nov., L. haddadi   sp. nov., and L. sofiae   sp. nov.). Palpal tibia with abundant rigid setae proventrally ( Fig. 3E View FIGURE , 5E View FIGURE , 9E View FIGURE , 12E View FIGURE , 14E View FIGURE , 21E View FIGURE ) and thin setae dorsally (thick in L. dippenaarae   sp. nov. and L. rattrayi   , Fig. 14D View FIGURE , 21D View FIGURE ). Males of L. engelbrechti   sp. nov., L. haddadi   sp. nov., and L. sofiae   sp. nov. with the palp tibiae strongly incrassate, ventrally excavated and with the posterior part of the ventral excavation produced ( Fig. 17D, E View FIGURE ; 19E, F View FIGURE ; 23D, E View FIGURE ). Males with a short ( Fig. 3I View FIGURE ) or elongated cuticular spur ( Figs. 5I, J View FIGURE , 6C, D View FIGURE , 9I, J View FIGURE , 12I, J View FIGURE , 14I View FIGURE , 21I View FIGURE ), with an apical megaespine on tibiae I. Tibiae I of males can be swollen (Group haddadi) or not (Group dregei). Males of L. engelbrechti   sp. nov., L. haddadi   sp. nov., and L. sofiae   sp. nov. present a strong spine near the base of the megaspine ( Fig. 17J View FIGURE ; 19K View FIGURE ; 23J View FIGURE ).

Metatarsus I is usually elongate and straight (Group dregei) or short and slightly bowed (Group haddadi). In the medial-basal part of the segment of all species (except L. confusus   sp. nov.) a prolateral protuberance (knob) is present and covered with short, black setae (Group Dregei, Figs. 5J View FIGURE , 9J View FIGURE , 12J View FIGURE ) or in the 1/3 A of the segment covered by black and blunt spinules (Group haddadi, Figs. 14I, J View FIGURE , 17I View FIGURE , 19J View FIGURE , 21I View FIGURE , 23I View FIGURE ). Both sexes with two rows of teeth on all paired claws; third claw short, curved. Scopulae on metatarsi I-II of females dense (sparse in males) and uniformly distributed along the length of the segment (sometimes with scattered, long, thin, emergent setae); III–IV sparse (1:2 A, sometimes divided) or absent. Tarsi I–II (both sexes) with dense scopulae, uniformly distributed throughout the segment; III–IV, less dense than I–II, uniformly distributed, and divided by band of setae. Tarsal organ low, tarsi integral. Trichobothria filiform; tibia in two rows; metatarsi with one straight line, tarsi with a zigzag row along full length. Metatarsal preening combs present in both sexes on legs II–IV (e.g. Fig. 2A View FIGURE ).

Copulatory bulb pyriform, embolus relatively short and tapering with lateral keels ( Figs. 3F, G View FIGURE , 9G, H View FIGURE , 12G View FIGURE , 14 View FIGURE F–H, 21F, G) or parallel ridges ( Figs. 3F, H View FIGURE , 5F, H View FIGURE ). In the species L. engelbrechti   sp. nov., L. haddadi   sp. nov., and L. sofiae   sp. nov. the copulatory bulb is generally smooth, with an embolus long and slightly twisted (keels present only in L. sofiae   sp. nov., Fig. 23 View FIGURE G-H). Females with two separate and small spermathecae, with a small base, a thin stalk, and an oval to rounded receptaculum ( Figs. 4E View FIGURE , 7C View FIGURE , 8E View FIGURE , 10D View FIGURE , 11D, 13D, 15D, 22D) or tubular cruved and without stalk ( Figs. 18D View FIGURE , 20D View FIGURE ). Apical segment of PLS elongated.

Distribution. Western Cape, Eastern Cape, Kwazulu-Natal, Free State and Mpumalanga Provinces, South Africa ( Fig. 24 View FIGURE , 25 View FIGURE ).

Species included. Lepthercus confusus   sp. nov., Lepthercus dippenaarae   sp. nov., Lepthercus dregei Purcell, 1902   , Lepthercus engelbrechti   sp. nov., Lepthercus filmeri   sp. nov., Lepthercus haddadi   sp. nov., Lepthercus kwazuluensis   sp. nov., Lepthercus lawrencei   sp. nov., Lepthercus mandelai   sp. nov., Lepthercus rattrayi Hewitt, 1917   , Lepthercus sofiae   sp. nov.

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Nemesiidae

Loc

Lepthercus

Ríos-Tamayo, Duniesky & Lyle, Robin 2020
2020
Loc

Lepthercus

Raven, R. J. 1985: 86
Hewitt, J. 1917: 699
Purcell, W. F. 1902: 379
1902