Rudolfina pilosa, Paiero & Marshall, 2020

Paiero, Steven Mark & Marshall, Stephen A., 2020, A revision of the genus Rudolfina Roháček (Sphaeroceridae: Limosininae), European Journal of Taxonomy 593, pp. 1-48 : 38-40

Rudolfina pilosa sp. nov.

urn:lsid:zoobank.org:act:8E664032-9FD5-4ED0-9A39-B63395A39655

Figs 18 , 21C

Etymology

The species epithet is from the Latin for ‘hairy’, referring to the hirsute sternites 4 and 5 of the male.

Material examined

Holotype

MEXICO • ♂; Oaxaca, 2.0 mi. W of Jct. Mex. 175- Yuvila Rd ; 9500 ft a.s.l.; 8–19 Aug. 1973; A. Newton leg.; oak-pine, dung; FMNH debu01086240.

Paratypes

MEXICO – Oaxaca • 1 ♂; 6.6 mi. N of Ixtlan de Juarez ; 8300 ft a.s.l.; 10–18 Aug. 1973; A. Newton leg.; oak woodland, human dung; DEBU • 1 ♂; 4.1 mi. W of Jct. Mex. 175- Yuvila Rd ; 9300 ft a.s.l.; 8–19 Aug. 1973; A. Newton leg.; oak-fir-pine forest; UNAM • 1 ♂; same collection data as for preceding; oak-fir-pine forest; FMNH .

Description

BODY. Length: 1.5–1.8 mm. Eye height 2.5 × genal height. Head with 3 interfrontal setae and 5–6 small setae along inner margin of orbital plate. Gena with 2 strong setae and 7–9 smaller setulae. Acrostichal setulae in 4–6 rows. Costagial seta extending slightly beyond humeral break; second costal sector equal to third sector; length of M 1 between crossveins dm-cu and r-m 2.0× dm-cu; CuA 1 stub vein ~5.0 × M 1 stub vein. Male mid femur with 4–5 small setae in ventrobasal cluster. Male mid tibia with ventral comb of 7–8 robust setae on apical ½, midventral seta absent (female unknown).

MALE ABDOMEN ( Fig. 18 ). Sternite 3 and 4 with dense cluster of setae posteromedially. Posterior margin of sternite 5 with round teeth on each side of a medial emargination; emargination less than 1/6 width of sternite and extending ~½ length. Transverse part of sternite 6 weakly arcuate. Epandrium with one large pair of setae ventrolaterally, one smaller pair of setae lateral to larger pair, and several setulae scattered over surface; subanal plate broadly continuous below anal opening. Surstylus boot-shaped, posteriorly bent near base, hirsute with 3–5 setae on posterior lobe (heel) and numerous smaller setae on anterior lobe (toe). Cercus fused with epandrium, with distinct seta on ventral margin. Postgonite clavate, with several setulae on dorsal surface. Dorsal sclerite of distiphallus ( Fig. 18 D–F) without distinct swellings and not extending beyond apex of acrophallus.

Female

Unknown.

Remarks

This species is most similar to R. newtoni sp. nov., which also has a setulose male sternite 4 and sternite 5. These species are easily distinguished by the shape of the male surstylus.

Figures

Fig. 18. Rudolfina pilosa sp. nov., male terminalia (debu01086241). A. Epandrium, surstylus and cercus, caudal view. B. Same as preceding, lateral view. C. Sternite 5, ventral view. D. Phallus and ejaculatory apodeme, dorsal view. E. Same as preceding, dorsolateral view. F. Same as preceding, lateral view. G. Postgonite, lateral view. Scale bars: A–B = 0.10 mm; C = 0.10 mm; D–G = 0.10 mm.

Fig. 21. Distribution of species of New World Rudolfina Roháček, 1987. A. R. cavernicola Marshall & Fitzgerald, 1997, R. digitata Marshall, 1991 and R. tumida sp. nov. B. R. bucki sp. nov., R. megepandria sp. nov. and R. howdeni sp. nov. C. R. pauca sp. nov., R. pilosa sp. nov., R. newtoni sp. nov. and R. remiforma sp. nov. D. R. exuberata sp. nov.