Lomachaeta, Mickel, 1936

Williams, Kevin A., Cambra, Roberto A., Bartholomay, Pedro R., Luz, David R., Quintero, Diomedes & Pitts, James P., 2019, Review of the genus Lomachaeta Mickel, 1936 (Hymenoptera: Mutillidae) with new species and sex associations, Zootaxa 4564 (1), pp. 101-136: 104-105

publication ID

https://doi.org/10.11646/zootaxa.4564.1.4

publication LSID

lsid:zoobank.org:pub:D0A9801B-8049-4211-A4A7-D7792B9D6936

persistent identifier

http://treatment.plazi.org/id/03F187A0-7C05-CD37-27B7-9DB7FA79FA11

treatment provided by

Plazi

scientific name

Lomachaeta
status

 

Key to species of Lomachaeta   (Males)

1. Ventral margin of mandible armed with strong tooth basally................................................... 2

- Ventral margin of mandible lacking tooth basally............................................................ 4

2(1). Femora and tegulae orange brown, contrasting with entirely black mesosoma and metasoma ( Fig. 42 View FIGURES 41–46 ; southwestern USA and northwestern Mexico)..................................................... L. cirrhomeris Pitts & Manley, 2004  

- Femora and tegula usually blackish, mesosoma blackish, metasoma often partly reddened; if femora orange, then metasoma also predominantly orange ( Figs 44–45 View FIGURES 41–46 )................................................................... 3

3(2). Penis valve with one basal tooth (see Pitts & Manley 2004, Fig. 16–18 View FIGURES 15–20 ); body color variable, entirely black or with metasoma cuticle variably orange to red ( Figs 43–46 View FIGURES 41–46 ; widespread in Nearctic)............................ L. hicksi Mickel, 1936  

- Penis valve with two basal teeth (see Pitts & Manley 2004, Fig. 14 View FIGURES 9–14 ); body entirely black ( Fig. 42 View FIGURES 41–46 ; Nayarit, Mexico South to Costa Rica)............................................................. L. chionothrix Pitts & Manley, 2004  

4(1). Metasoma largely red to orange (e.g. Figs 47–52 View FIGURES 47–52 ; North America only).......................................... 5

- Body entirely black except tegula, head, or legs sometimes orange or lighter brown ( Figs 53–64 View FIGURES 53–56 View FIGURES 57–60 View FIGURES 61–64 ; North or South America). 9

5(4). Paramere broadly flattened throughout, rounded apically (Arizona, California, and Nevada; Fig. 47 View FIGURES 47–52 , also see Pitts & Manley 2004, Fig. 23 View FIGURES 21–26 ).......................................................... L. beadugrimi ( Pitts & Manley 2004)  

- Paramere acuminate apically, at most laterally compressed (e.g. Pitts & Manley 2004, Figs 14–22 View FIGURES 9–14 View FIGURES 15–20 View FIGURES 21–26 ).................... 6

6(5). Wing venation reduced, cells contained in basal half of forewing ( Figs 48, 49 View FIGURES 47–52 ); paramere lacking row of elongate setae ventrally (e.g. Fig. 67 View FIGURES 65–72 ).................................................................................... 7

- Wing venation normal, cells extending beyond basal half of forewing (e.g. Fig. 52 View FIGURES 47–52 ); paramere with somewhat uniform row of long setae ventrally (see Manley & Pitts 2004; Fig. 19, 20 View FIGURES 15–20 )................................................... 8

7(1). Head and mesosoma orange, concolorous with T2 ( Fig. 49 View FIGURES 47–52 ); paramere laterally compressed, moderately setose ( Fig. 49, 50 View FIGURES 47–52 ; Riverside County, California)................................................. L. calamondin Williams   , sp. nov.

- Head and mesosoma black ( Fig. 48 View FIGURES 47–52 ); paramere not so compressed, basically subcylindrical, sparsely setose (Central Valley and Coast Ranges, California)....................................................... L. powelli ( Mickel), 1964  

8(7). Pronotum and mesoscutum with IS more than twice PD; metasoma blackish with orange color restricted to T2 and T3 ( Fig. 51 View FIGURES 47–52 ; Sonoran Desert in Arizona and California, USA and Oaxaca, Mexico)............ L. ptilohyalus Pitts & Manley, 2004  

- Pronotum and mesoscutum with IS less than twice PD; metasoma mostly orange ( Fig. 52 View FIGURES 47–52 ; Great Basin to Chihuahuan Desert in southwestern Nearctic Region)............................................ L. crocopinna Pitts & Manley, 2004  

9(7). Apical fringe of T2 with row of convergent thick bristles (e.g. Figs 53, 54 View FIGURES 53–56 , 57 View FIGURES 57–60 ; North and South America).............. 10

- Apical fringe of T2 with simple setae only (e.g. Figs 58, 60 View FIGURES 57–60 , 63 View FIGURES 61–64 ; North America).................................. 13

10(9). Vertex with separated punctures, IS=PD; paramere having long setae on external surface medially (see Williams & Pitts 20 0 9, Fig. 15 View FIGURES 15–20 ; Sonora, Mexico)..................................................... L. theresa Williams & Pitts, 2009  

- Frons and vertex coarsely punctate or areolate; paramere lacking long setae except sometimes at apex (South America)... 11

11(10). T1 shape nodose, anterior face convex ( Fig. 56 View FIGURES 53–56 ); apical third of paramere distinctly downcurved ( Fig. 71 View FIGURES 65–72 ; Colombia).................................................................................... L. juanita Williams   , sp. nov.

- T1 shape disciform, anterior face flat or concave ( Figs 53, 55 View FIGURES 53–56 ); paramere basically straight or evenly downcurving throughout its length (e.g. Manley & Pitts 2004, Fig. 21 View FIGURES 21–26 ).............................................................. 12

12(11). Gena weakly carinate, apex of paramere with weak tuft of long setae (see Williams & Pitts 2009, Fig. 7 View FIGURES 5–8 ; Argentina, Brazil, and Paraguay)....................................................................... L. vianai Casal, 1969  

- Genal carina distinct; apex of paramere lacking setal tuft (see Manley & Pitts 2004, Fig. 21 View FIGURES 21–26 ; Northern South America)............................................................................. L. hyphantria Pitts & Manley, 2004  

13(9). Paramere broadly flattened, rounded apically ( Fig. 69 View FIGURES 65–72 ; also see Williams & Pitts 2009, Fig. 13 View FIGURES 9–14 )..................... 14

- Paramere acuminate apically, at most laterally compressed (e.g. see Williams & Pitts 2009, Fig. 1 View FIGURES 1–4 )................... 15

14(13). Cuspis and paramere with all setae shorter than paramere width (see Williams & Pitts 2009, Fig. 13 View FIGURES 9–14 ; Sonoran Desert in California)................................................................ L. snellingella Williams & Pitts, 2009  

- Cuspis and paramere with setae much longer than paramere width ( Figs 59 View FIGURES 57–60 , 69 View FIGURES 65–72 ; Sonora, Mexico)................................................................................................... L. eironeia Williams   , sp. nov.

15(13). Paramere lacking long setae, all setae shorter than paramere width (see Williams & Pitts 2009, Fig. 9 View FIGURES 9–14 ; Arizona, USA and Sonora, Mexico).................................................... L. polemomechana Williams & Pitts, 2009  

- Paramere with at least some ventral or apical setae longer than mid-paramere width............................... 16

16(15). Entire paramere with continuous ventral setal row (see Williams & Pitts 2009, Fig. 3 View FIGURES 1–4 ; Mohave and Sonoran Deserts in California and Nevada)............................................................... L. ilex Williams & Pitts, 2009  

- Long ventral setae of paramere absent from basal half....................................................... 17

17(16). Paramere narrow throughout, long cylindrical, apically convergent, and evenly curving ventrally throughout length ( Fig. 63 View FIGURES 61–64 , also see Williams & Pitts 2009, Fig. 5 View FIGURES 5–8 ; Arizona, USA and Sonora, Mexico)........... L. litosisyra Williams & Pitts, 2009  

- Paramere evenly tapering toward apex, somewhat laterally compressed, subparallel, and virtually straight to apex or scarcely downcurving (e.g. Figs 62, 64 View FIGURES 61–64 )......................................................................... 18

18(17). Long setae of paramere restricted to tuft in apical fifth of free length (see Williams & Pitts 2009, Fig. 17 View FIGURES 15–20 ; Texas, New Mexico, and Arizona, USA and Sonora, Mexico)..................................... L. vacamuerta Williams & Pitts, 2009  

- Long setae of paramere scattered through apical half of free length (see Williams & Pitts 2009, Fig. 1 View FIGURES 1–4 ; Baja California Sur, Mexico).................................................................. L. hedera Williams & Pitts, 2009