Martiodrilus (Maipure) Righi, 1995

Feijoo, Alexander, Zuluaga, Luis Fernando & Molina, Ligia Janneth, 2018, New species and records of earthworms (Annelida, Oligochaeta) in plantain cropping systems in Colombia's coffee-growing region, Zootaxa 4496 (1), pp. 448-458 : 452

Subgenus Martiodrilus (Maipure) Righi, 1995

Martiodrilus (Maipure) Righi, 1995: 531 . Zicsi 2001: 114; 2005: 113.

Type species. Rhinodrilus ecuadoriensis Benham, 1892 .

Diagnosis. Number of species: 15. Distribution: Colombia (6), Ecuador (4), Peru (2), Guyana (2), Surinan (1). In some cases, the species are distributed in several countries, for example M. (Mp.) agricola , M. (Mp.) ecuadoriensis , and M. (Mp.) savanicola , in Colombia and Ecuador and M. (Mp.) micrurus and M. (Mp.) ecuadoriensis in Ecuador and Peru ( Feijoo& Celis 2011).

Genus diagnosis, modified, following Righi (1995), Zicsi (2001): Species with green, red, violet, brown or blueish pigmentation. Eight rows of regular, closely paired setae. Absent or very thin membranous septa in the gizzard area; 7–8 pairs of calciferous glands between segments 7–13 or 8–14 with a composite, reticulate tubular structure. Holandric and metagynic sexual apparatus with testes in segments 10 and 11 and hypo- or periesophageal testis sacs. Short seminal vesicles limited to segments in 11 and 12. Four pairs of spermathecae present in segments 6, 7, 8, 9.