Syzeuctus fuscator ( PANZER , 1809)

Riedel, Matthias, 2021, The genus Syzeuctus FÖRSTER in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species, Linzer biologische Beiträge 52 (2), pp. 1013-1098 : 1042-1043

Syzeuctus fuscator ( PANZER, 1809) (Figs 71, 114)

O r i g i n a l: Ophion fuscator PANZER, 1809 : vol 102, fig. 22. Type lost, interpretation according to

HORSTMANN 1982: 236. syn. Syzeuctus buculus SEYRIG, 1927 , synonymized by HORSTMANN 1999: 53. L e c t o t y p e: (♀) "Htes Alpes 5.26-Cleu" (Munich), type studied. syn. Syzeuctus hispanicus HEDWIG, 1939 , synonymized by HORSTMANN 1999: 53. Holotype from

Valencia / Spain lost. S t u d i e d m a t e r i a l: 51♀♀, 51♂♂ ; Czech Republic (Moravia)*, France, Germany ,

Hungary, Italy *, Mallorca, Morocco, Portugal, Sardinia*, Sicily*, Spain, Switzerland *, United

Kingdom. D e s c r i p t i o n: ♀: Body length 10-12 mm. Antenna with 33-39 flagellomeres; 1st

flagellomere 4.6-5.0× longer than wide and 0.55× as long as eye; preapical flagellomere about square. Head 2.2× wider than long. Temple strongly and roundly narrowed behind eye, dorsally 0.35× as long as eye. OED 1.6-2.2× ocellar diameter. Frons ± impressed medially, slightly or strongly swollen laterally, with dense punctures. Face densely punctate, centrally protruding, inner eye margins slightly convergent ventrally. Clypeus c. 2× wider than long, with rather dense punctures, apical margin rounded. MI 0.5-0.7. Genal carina reaching mandibular base.

Mesosoma densely punctate, 1.45× longer than high. Frontoventral edge of pronotum moderately to strongly curved; lateral pronotal groove smooth. Speculum punctate. Epicnemial carina usually moderately elevated ventrally, with dentiform lamella on transversal part. Apical transversal carina of propodeum complete. Hind femur 4.8-6.0× longer than wide. Hind metatarsus 0.65× as long as hind femur. 2 nd hind tarsomere 1.5× longer than 3 rd one. Hind claw with short basal teeth, moderately bent apically. Areolet stalked, 2 nd recurrent vein in or slightly distal to its middle. Nervulus postfurcal by 1-2× its width. Hairs on basal cell about as long as their intervals. Hind wing with 9-10 distal hamuli.

1 st tergite 1.5-2.1× longer than wide, with scattered lateral punctures. 2 nd tergite 0.9- 1.25× as long as wide, with dense punctures. 3 rd tergite 0.9-1.2× as long as wide, densely punctate. Spiracles of 1 st to 3 rd tergites ± protruding laterally. Ovipositor 1.0-1.4× as long as fore wing, 1.1-1.4× longer than metasoma and 2.1-3.0× longer than hind tibia.

Color: Black. Clypeus, sometimes mandibular base, wider inner and narrow outer orbits, frontal spot on upper margin of pronotum, anterolateral and central spots on mesoscutum, spot on mesopleuron, tegula, spot on subtegular ridge, scutellum except anteromedian black spot, postscutellum and apical band on propodeum yellow. Metasoma black; 1 st tergite often with subbasal yellow band; sometimes 2 nd and 3 rd tergites with basal yellow bands; 1 st to 3 rd tergites with wide apical yellow bands (sometimes also 4 th tergite). Coxae and mid and hind trochanters black, sometimes fore coxa with yellow spot; fore and mid legs yellow or reddish-yellow; hind femur black, brown or reddish-brown (but always darker than mid femur), often yellow apically; hind tibia yellow, brownish or black in apical third, hind tarsus ± reddish or brownish, sometimes (in a specimen from Morocco) hind metatarsus with central yellow ring. Wings with apical brown stripes (often weak); pterostigma reddish-brown, sometimes paler proximally.

♂: Body length 8-12 mm. Antenna with 33-40 flagellomeres; 1 st flagellomere 4.2× longer than wide. Temple moderately and roundly narrowed behind eye. OED 1.7-2.0× ocellar diameter. MI 0.6-0.8. Epicnemial carina slightly elevated, with dentiform lamella at transversal part. Apical transversal carina of propodeum complete. Hind femur 4.8- 5.9× longer than wide. 1 st tergite 1.9-2.0× longer than wide, with rather coarse but scattered punctures. 2 nd tergite 1.0-1.2× longer than wide, with coarse rather dense punctures. 3 rd tergite 1.0-1.2× longer than wide, with coarse dense but sometimes superficial punctures.

Color: Black. Face ± yellow, sometimes with black median stripes. Palps, mandible, clypeus, malar space, and wide orbits yellow. Sometimes scape, pedicel, and flagellum yellowish ventrally. Mesosoma with extended yellow coloration. 1 st to 4 th tergites with apical yellow bands (often also with yellow basal spots or bands), rarely 5 th tergite with weak ochreous apical band. Fore and mid legs except black bases of coxae completely yellow. Hind femur dark brown or black, sometimes yellow in apical 0.1; hind tibia yellow, infuscate in apical 0.4; hind tarsus reddish-yellow or brown. Wings sometimes ± infuscate, with weak brown apical bands; pterostigma ochreous.

Hosts: Epischnia asteris (STAUDINGER, 1870) (Lep. Pyralidae ) (Edinburgh), Pempelia palumbella (DENIS & SCHIFFERMÜLLER, 1775) (Lep. Pyralidae ) (Edinburgh), Oiceticoides febretta (FONSCOLOMBE, 1835) (Lep. Psychidae ) ( AUBERT 1978: 126).