Syzeuctus exsculptus ( FONSCOLOMBE , 1854)

Riedel, Matthias, 2021, The genus Syzeuctus FÖRSTER in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species, Linzer biologische Beiträge 52 (2), pp. 1013-1098 : 1036-1037

Syzeuctus exsculptus ( FONSCOLOMBE, 1854) (Figs 67, 110)

O r i g i n a l: Lissonota exsculpta FONSCOLOMBE, 1854: 509

H o l o t y p e: (♀) "18. L. exsculpta nob." (Paris), type not studied.

syn. Lissonota pubescens FONSCOLOMBE, 1854: 514 , synonymized by AUBERT 1978: 128

H o l o t y p e: (♂)"26. L. pubescens nob." (Paris), type not studied.

syn. nov. Lissonota puberula KRIECHBAUMER, 1895: 263-264

H o l o t y p e: (♂) (Bern), type not studied.

syn. nov. Syzeuctus schmiedeknechti BRAUNS, 1901: 182 , synonymized by AUBERT 1969: 87.

L e c t o t y p e: (♀) " Brauns det.; 9658; 17. VI.89 Berisal " (Bern), not studied .

P a r a l e c t o t y p e: (♂) "17.VI.89 Berisal" (Berlin), studied.

S t u d i e d m a t e r i a l: 41♀♀, 40♂♂; Bulgaria *, France, Mongolia *, Morocco *, Spain, Switzerland, Turkey.

D e s c r i p t i o n: ♀: Body length 9-12 mm. Antenna with 40-45 flagellomeres; 1 st flagellomere 4.0-5.0× longer than wide and 0.52-0.65× as long as eye; preapical flagellomere c. 1.2× longer than wide. Temple shortly parallel or slightly narrowed behind eye, moderately and roundly narrowed apically, dorsally 0.4-0.45× as long as eye. Ocelli small, OED 2.0-2.6× ocellar diameter. Frons punctate, basally often smooth, slightly impressed medially. Clypeus 1.5-1.8× wider than long, apex rounded. MI 0.5- 0.7. Genal carina reaching mandibular base.

Mesosoma 1.3-1.35× longer than high. Ventrofrontal edge of pronotum moderately curved; lateral pronotal groove rugose-punctate or with fine striae. Epicnemial carina often moderately elevated, with ± dentiform or sometimes low lamella at transversal part. Propodeum with complete apical transversal carina, carina sometimes shortly indistinct medially. Hind femur 5.0-5.5× longer than wide. Hind metatarsus c. 0.6× as long as hind femur. 2 nd hind tarsomere c. 1.5× longer than 3 rd one. Hind claw completely pectinate, moderately curved apically. Areolet stalked; 2 nd recurrent vein in or slightly proximal or distal to its middle. Nervulus interstitial or slightly postfurcal by 1× its width. Hairs of basal cell as long as their intervals. Hind wing with 8-9 distal hamuli.

Metasoma rather stout. 1 st tergite 1.55-1.9× longer than wide, with scattered or rather dense, coarse punctures laterally. 2 nd tergite 0.9-1.1× as long as wide, with rather dense coarse punctures. 3 rd tergite 0.9-1.1× as long as wide, with dense but sometimes superficial punctures. Ovipositor sheath 1.0-1.4× longer than fore wing, 1.2-1.5× longer than metasoma and 2.3-2.9× longer than hind tibia.

Color: Black. Outer orbit up to vertex with narrow yellow stripe. Clypeus often ±

reddish. Mesosoma black. 1 st to 3 rd tergites red; sometimes 1 st tergite black basally and 4 th tergites red basally. Coxae and trochanters black; legs black, with ± extended yellow frontal stripes on fore and mid femora, tibiae and tarsi; or fore and mid legs and hind femur reddish-brown. Wings ± infuscate, sometimes with weak brown apical spot;

pterostigma dark brown. ♂: Body length 8.5-12 mm. Antenna with 40-45 flagellomeres; 1st flagellomere 4.5×

longer than wide; preapical flagellomere 1.0-1.1× longer than wide. Head and mesosoma with long erected hairs. Temple shortly parallel behind eye, slightly and roundly narrowed apically, dorsally 0.5-0.6× as long as eye. OED 1.7-2.5× ocellar diameter. MI 0.7. Genal carina reaching mandibular base.

Mesosoma with long erected hairs. Lateral pronotal groove rugose or punctate. Speculum punctate. Epicnemial carina moderately elevated, with dentiform lamella on transversal part. Apical transversal carina complete. Hind femur 4.4-5.5× (in one ♂ from Morocco 6.3×) longer than wide. Hind metatarsus 0.7× as long as hind femur. 2 nd hind tarsomere 1.35× longer than 3 rd one. Hind claw with basal teeth, moderately bent apically. Areolet stalked; 2 nd recurrent vein in its middle. Nervulus interstitial to postfurcal by 2× its width. Hairs on basal cell as long as their intervals. Hind wing with 8 distal hamuli.

1 st tergite 1.5-1.9× longer than wide, with scattered to rather dense punctures. 2 nd tergite 1.0-1.1× wider than long, with dense punctures. 3 rd tergite 1.0-1.2× wider than long, with dense punctures.

Color: Black. Clypeus blackish or reddish. Outer orbit and sometimes vertex narrowly yellow. 1 st tergite black, sometimes red apically; 2 nd to 4 th /5 th tergites completely red. Coxae and trochanters black. Legs otherwise blackish or ± reddish; fore and mid femora blackish basally and yellowish apically, fore and mid tibiae and tarsi yellowish frontally; hind femur reddish, brown or black; hind tibia often with yellow base, sometimes with external yellow stripe; hind metatarsus and 2 nd hind tarsomere ± yellow basally. Wings

without brown spots, pterostigma brown. One ♂ from Morocco differs by: slenderer hind femur (6.3× longer than wide), red 5th

tergite and completely black hind tarsus.

R e m a r k: In a long series of ♀♀ and ♂♂ from Monetier-les Bains/ France collected by R. HINZ (material in ZSM, Munich), hind femora of the specimens were red or black without other differences of structure or coloration. AUBERT (1978: 132) noted that Lissonota puberula differs from S. exsculptus by a stouter hind femur and yellow-red clypeus. Both features are variable in this species. I therefore synonymize both species here.