Paradilacra densissima ( Bernhauer, 1909 )

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134: 24-29

publication ID

http://doi.org/ 10.11646/zootaxa.353.1.1

publication LSID

lsid:zoobank.org:pub:104E9C0B-B207-41E6-816C-E91FC1C7DBA0

DOI

http://doi.org/10.5281/zenodo.5102678

persistent identifier

http://treatment.plazi.org/id/03F387B2-FF97-5350-CD24-528BFDA7A3B9

treatment provided by

Felipe

scientific name

Paradilacra densissima ( Bernhauer, 1909 )
status

 

Paradilacra densissima ( Bernhauer, 1909)  

( Figs. 19–30 View FIGURES 19–22 View FIGURES 23–30 )

Atheta (Paradilacra) densissima Bernhauer, 1909: 517   .

Paradilacra persola Casey, 1910a: 72   .

Paradilacra willametta Casey, 1910a: 73   .

Paradilacra uintana Casey, 1910a: 73   .

Paradilacra glenorica Casey, 1910a: 74   .

Paradilacra symbolica Casey, 1911a: 127   .

Paradilacra erebea Casey, 1911a: 128   .

Paradilacra subaequa Casey, 1911a: 128   .

Paradilacra sinistra Casey, 1911a: 129   .

Paradilacra memnonia Casey, 1911a: 130   .

Paradilacra vulgatula Casey, 1911a: 130   .

Paradilacra deserticola Casey, 1911a: 131   .

Paradilacra densissima: Fenyes, 1920: 244   (as valid species).

Paradilacra persola: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra willametta: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra uintana: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra glenorica: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra symbolica: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra erebea: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra subaequa: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra sinistra: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra memnonia: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra vulgatula: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Paradilacra deserticola: Fenyes, 1920: 244   (as synonym of Pa. densissima   ).

Atheta (Paradilacra) densissima: Bernhauer & Scheerpeltz, 1926: 607   (as valid species).

Atheta (Paradilacra) persola: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) willametta: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) uintana: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) glenorica: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) symbolica: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) erebea: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) subaequa: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) sinistra: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) memnonia: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima).

Atheta (Paradilacra) vulgatula: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) deserticola: Bernhauer & Scheerpeltz, 1926: 607   (as synonym of At. densissima   ).

Atheta (Paradilacra) densissima: Moore & Legner, 1975: 358   (as valid species).

Atheta (Paradilacra) persola: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Atheta (Paradilacra) willametta: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Atheta (Paradilacra) uintana: Moore & Legner, 1975: 358   (as synonym of At. densissima   ; misspelled as uinata).

Atheta (Paradilacra) glenorica: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Atheta (Paradilacra) symbolica: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Atheta (Paradilacra) erebea: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Atheta (Paradilacra) subaequa: Moore & Legner, 1975: 358   (as synonym of At. densissima   ; misspelled as subaegua).

Atheta (Paradilacra) sinistra: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Atheta (Paradilacra) memnonia: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Atheta (Paradilacra) vulgatula: Moore & Legner, 1975: 358   (as synonym of At. densissima   ; misspelled as vulgata).

Atheta (Paradilacra) deserticola: Moore & Legner, 1975: 358   (as synonym of At. densissima   ).

Paradilacra densissima: Seevers, 1978: 265   (as valid species).

Paradilacra persola: Seevers, 1978: 265   (as valid species).

Paradilacra willametta: Seevers, 1978: 266   (as valid species).

Paradilacra uintana: Seevers, 1978: 266   (as valid species).

Paradilacra glenorica: Seevers, 1978: 265   (as valid species).

Paradilacra symbolica: Seevers, 1978: 265   (as valid species).

Paradilacra erebea: Seevers, 1978: 265   (as valid species).

Paradilacra subaequa: Seevers, 1978: 265   (as valid species).

Paradilacra sinistra: Seevers, 1978: 265   (as valid species).

Paradilacra memnonia: Seevers, 1978: 265   (as valid species).

Paradilacra vulgatula: Seevers, 1978: 266   (as valid species).

Paradilacra deserticola: Seevers, 1978: 265   (as valid species).

Type material. Lectotype of Atheta densissima   (here designated): ♂, “ S. Anselmo Cal. [ifornia]”, “June 04.16 [4.vi.1916]”, “166.”, “ ambigua Er.   ? Fenyes. det. Bernhauer ”, “ densissima Bernh.   Typus. Fenyes” (yellow label), “not ambigua Erichson   det. Lohse ”, ( FMNH)   . Paralectotype: ♂, “Willist’n [Williston], N.[orth] D.[akota] Jun.[e] 8–9 Wickham”, “7614”, “ densissima Bernh. Cotypus. Fenyes   ” (yellow label) ( FMNH)   .

Lectotype of Paradilacra persola   (here designated): ♂, “Cal.[ifornia, Monterey]”, “ Paradilacra persola Csy.   ”, “TYPE USNM 39377” (red label), “CASEY bequest 1925” ( NMNH). Paralectotype: ♂, “Cal.[ifornia, Monterey]”, “ persola   PARATYPE USNM 39377” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Paradilacra willametta   (here designated): ♂, “Portland Oreg.[on]”, “ willametta Csy.   ”, “TYPE USNM 39380” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Paradilacra uintana   (here designated): ♂, “Milford Ut.[ah] Jul.[y] Wickha[m]”, “ uintana Csy.   ”, “TYPE USNM 39379” (red label), “CASEY bequest 1925” ( NMNH). Paralectotypes: ♂, “Milford Ut.[ah] Jul.[y] Wickha[m]”, “ uintana   ­2 PARATYPE USNM 39379” (red label), “CASEY bequest 1925”; ♂, “Elko, Nevad[a] Wickham.”, “ uintana   ­3 PARATYPE USNM 39379” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Paradilacra glenorica   (here designated): ♂, “Glenora B.[ritish] C.[olumbia]”, “ glenorica Csy.   ”, “TYPE USNM 39378” (red label), “CASEY bequest 1925” ( NMNH). Paralectotype: ♀, “Kamloops, B.[ritish] C.[olumbia] Wickham.”, “ glenorica   ­3 PARATYPE USNM 39378” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Paradilacra symbolica   (here designated): ♀, “Cal.[ifornia, Napa Junction]”, “ symbolica Csy.   ”, “TYPE USNM 39385” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Paradilacra erebea   (here designated): ♀, “Cal.[ifornia, Napa Junction]”, “ erebea Csy.   ”, “TYPE USNM 39381” (red label), “CASEY bequest 1925”. Paralectotype: ♂, “Cal.[ifornia, near Mt. Diablo]”, “ erebea   ­2 PARATYPE USNM 39381” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Paradilacra subaequa   (here designated): ♂, “Cal.[ifornia, Santa Rosa]”, “ subaequa Csy.   ”, “TYPE USNM 39382” (red label), “CASEY bequest 1925”. Paralectotype: ♂, “Cal.[ifornia, Santa Rosa]”, “ subaequa   ­2 PARATYPE USNM 39382” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Paradilacra sinistra   (here designated): ♂, “Cal.[ifornia, San José and Santa Clara Valley]”, “ sinistra Csy.   ”, “TYPE USNM 39383” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Paradilacra memnonia   (here designated): ♀, “Cal.[ifornia, Cloverdale , north of Santa Rosa]”, “ memnonia Csy.   ”, “TYPE USNM 39384” (red label), “CASEY bequest 1925” ( NMNH).  

Holotype of Paradilacra vulgatula   : ♂, “Cal.[ifornia, Jountville , Napa Co.]”, “ vulgatula Csy.   ”, “TYPE USNM 39386” (red label), “CASEY bequest 1925”   . Paratype: ♀, “Cal.[ifornia, Jountville , Napa Co.]”, “ vulgatula   ­2 PARATYPE USNM 39386” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Paradilacra deserticola   (here designated): ♀, “Elko Nev.[ada]”, “ deserticola Csy.   ”, “TYPE USNM 39387” (red label), “CASEY bequest 1925” ( NMNH).  

Additional material. UNITED STATES: California: ♀, “Cal” with a black dot inside “C” ( San Francisco and immediate vicinity as far south as Redwood City and Purissima ( FitzGerald 1962)) ( NMNH (Casey collection)); San Bernardino Co   .: 32 specimens, 4 km NEE Big Bear City, Hwy. 18, San Bernardino Nat. Forest, 34º16.46’N 116º48.07’W, 2050 m, Baldwin Lake bank, 11.viii.1999 (V.I.Gusarov) ( KSEM) GoogleMaps   ; San Diego Co   .: 3♂♂, 5♀♀, Cuyamaca , 11.vi.1961 (I.Moore) ( CNCI, SPSU)   ; San Luis Obispo Co.: ♂, Atascadero , 15.viii.1956 (I.Moore) ( CNCI)   ; Ventura Co.: 14 specimens, 10 km N Ojai, Los Padres Nat. Forest, Rose Valley Rd. , 34º32.51’N 119º11.42’W, 1150 m, reservoir bank, 10.viii.1999 (V.I.Gusarov) ( KSEM) GoogleMaps   ; Montana: ♂, Glendive (This specimen was labeled by curators as a paratype of Pa. glenorica   but it had not been included in the type series of that species by Casey (1910a) ( NMNH (Casey collection)); CANADA: Alberta: ♂, ♀, Waterton Lakes N. P., bes. Hwy. 6, 4.4 km SE jct. Hwy. 5, tread beaver pond, 6.vi.1980 ( CNCI)   ; British Columbia: ♂, Thompson River, Monte Creek , 27.vi.1988 (L.LeSage)   ; ♂, 16 mi. W Osoyoos , 5.vi.1968 (Campbell & Smetana) ( CNCI)   .

Diagnosis. Paradilacra densissima   is the only valid Nearctic species of the genus. When describing Paradilacra, Bernhauer (1909)   included in it an additional Nearctic species ( Atheta ambigua ( Erichson, 1839))   and one species from North Africa ( At. hyperbolica Bernhauer, 1905   ). In this paper At. ambigua   is placed in the genus Strigota Casey, 1910a   (see below). The aedeagus and spermatheca of Pa. densissima   are shown in Figs. 23–30 View FIGURES 23–30 .

Discussion. My comparison of the types of the species described by Casey (1910a, 1911a) in the genus Paradilacra   with the types of Pa. densissima   confirmed the synonymies established by Fenyes (1920). The body size and the length of the female spermatheca ( Figs. 29–30 View FIGURES 23–30 ) varies in examined specimens but this variation is continuous. In all examined male specimens the shape of the aedeagus and the sclerites of its internal sac were identical.

Distribution. Paradilacra densissima   is widely distributed in the western United States (California, Nevada, Oregon, Utah, Montana and North Dakota) and Canada (British Columbia and Alberta).

FMNH

Field Museum of Natural History

NMNH

Smithsonian Institution, National Museum of Natural History

CNCI

Canadian National Collection Insects

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Paradilacra

Loc

Paradilacra densissima ( Bernhauer, 1909 )

Gusarov, Vladimir I. 2003
2003
Loc

Paradilacra densissima: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra persola: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra willametta: Seevers, 1978: 266

Seevers, C. H. 1978: 266
1978
Loc

Paradilacra uintana: Seevers, 1978: 266

Seevers, C. H. 1978: 266
1978
Loc

Paradilacra glenorica: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra symbolica: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra erebea: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra subaequa: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra sinistra: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra memnonia: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Paradilacra vulgatula: Seevers, 1978: 266

Seevers, C. H. 1978: 266
1978
Loc

Paradilacra deserticola: Seevers, 1978: 265

Seevers, C. H. 1978: 265
1978
Loc

Atheta (Paradilacra) densissima: Moore & Legner, 1975: 358

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) persola:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) willametta:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) uintana:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) glenorica:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) symbolica:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) erebea:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) subaequa:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) sinistra:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) memnonia:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) vulgatula:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) deserticola:

Moore, I. & Legner, E. F. 1975: 358
1975
Loc

Atheta (Paradilacra) densissima: Bernhauer & Scheerpeltz, 1926: 607

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) persola:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) willametta:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) uintana:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) glenorica:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) symbolica:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) erebea:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) subaequa:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) sinistra:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) memnonia:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) vulgatula:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Atheta (Paradilacra) deserticola:

Bernhauer, M. & Scheerpeltz, O. 1926: 607
1926
Loc

Paradilacra densissima: Fenyes, 1920: 244

Fenyes, A. 1920: 244
1920
Loc

Paradilacra persola:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra willametta:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra uintana:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra glenorica:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra symbolica:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra erebea:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra subaequa:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra sinistra:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra memnonia:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra vulgatula:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra deserticola:

Fenyes, A. 1920: 244
1920
Loc

Paradilacra symbolica

Casey, T. L. 1911: 127
1911
Loc

Paradilacra erebea

Casey, T. L. 1911: 128
1911
Loc

Paradilacra subaequa

Casey, T. L. 1911: 128
1911
Loc

Paradilacra sinistra

Casey, T. L. 1911: 129
1911
Loc

Paradilacra memnonia

Casey, T. L. 1911: 130
1911
Loc

Paradilacra vulgatula

Casey, T. L. 1911: 130
1911
Loc

Paradilacra deserticola

Casey, T. L. 1911: 131
1911
Loc

Paradilacra persola

Casey, T. L. 1910: 72
1910
Loc

Paradilacra willametta

Casey, T. L. 1910: 73
1910
Loc

Paradilacra uintana

Casey, T. L. 1910: 73
1910
Loc

Paradilacra glenorica

Casey, T. L. 1910: 74
1910
Loc

Atheta (Paradilacra) densissima

Bernhauer, M. 1909: 517
1909