Atheta keeni Casey, 1910a, Casey, 1910

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134: 37-39

publication ID

http://doi.org/ 10.11646/zootaxa.353.1.1

publication LSID

lsid:zoobank.org:pub:104E9C0B-B207-41E6-816C-E91FC1C7DBA0

DOI

http://doi.org/10.5281/zenodo.5103048

persistent identifier

http://treatment.plazi.org/id/03F387B2-FFA4-536A-CD24-5076FB2FA7D9

treatment provided by

Felipe

scientific name

Atheta keeni Casey, 1910a
status

 

Atheta keeni Casey, 1910a  

( Fig. 97 View FIGURES 93–97 in Klimaszewski & Winchester 2002; Figs. 41–46 View FIGURES 41–46 in this paper)

Atheta (Lamiota) keeni Casey, 1910a: 17   .

Atheta (s. str.) innocens Casey, 1910a: 39   , syn. nov.

Atheta (Lamiota) achromata Casey, 1911a: 82   , syn. nov.

Atheta (Lamiota) profecta Casey, 1911a: 83   , syn. nov.

Atheta (Liogluta) keeni: Fenyes, 1920: 209   (as valid species).

Atheta (s. str.) innocens: Fenyes, 1920: 214   (as valid species).

Atheta (Liogluta) achromata: Fenyes, 1920: 209   (as valid species).

Atheta (Liogluta) profecta: Fenyes, 1920: 209   (as valid species).

Atheta (Liogluta) keeni: Bernhauer & Scheerpeltz, 1926: 657   (as valid species).

Atheta (s. str.) innocens: Bernhauer & Scheerpeltz, 1926: 644   (as valid species).

Atheta (Liogluta) achromata: Bernhauer & Scheerpeltz, 1926: 656   (as valid species).

Atheta (Liogluta) profecta: Bernhauer & Scheerpeltz, 1926: 658   (as valid species).

Atheta (Liogluta) keeni: Moore & Legner, 1975: 364   (as valid species).

Atheta (s. str.) innocens: Moore & Legner, 1975: 363   (as valid species).

Atheta (Liogluta) achromata: Moore & Legner, 1975: 353   (as valid species).

Atheta (Liogluta) profecta: Moore & Legner, 1975: 370   (as valid species).

Lamiota keeni: Seevers, 1978: 263   (as valid species).

Xenota innocens: Seevers, 1978: 270   (as valid species).

Lamiota achromata: Seevers, 1978: 263   (as valid species).

Lamiota profecta: Seevers, 1978: 263   (as valid species).

Atheta (Liogluta) vasta: Klimaszewski & Winchester, 2002: 39   (nec Lohse & Smetana, 1985: 297) (as valid species).

Type material. Lectotype of Atheta keeni   (here designated): ♂, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ Lamiota keeni Csy.   ”, “TYPE USNM 39480” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: 5♂♂, 5♀♀, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ keeni   PARATYPE USNM 39480” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta innocens   (here designated): ♀, “ Lane Co., Or.[egon]”, “ Atheta innocens Csy.   ”, “TYPE USNM 39270” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta achromata   (here designated): ♂ (without head and prothorax), “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ achromata Csy.   ”, “TYPE USNM 39481” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta profecta   (here designated): ♂, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ profecta Csy.   ”, “TYPE USNM 39482” (red label), “CASEY bequest 1925” ( NMNH).  

Paralectotype (see discussion) of Homalota vasta Mäklin   in Mannerheim, 1853: ♀, “Sitka”, “Holmberg”, “ Homalota vasta   m. Sitkha”, “Typus” (green label), “Mus. Zool. H:fors. Spec. Typ. No. 2250. Homalota vasta Mäklin   ”, “ Homalota vasta Mäkl.   Holotypus / Liogluta vasta   . Holotypus Lohse fix. 1985” ( MZHF)   .

Additional material. UNITED STATES: Alaska: ♀, Valdez , 61º09.05'N 146º21.80'W, 20 m, in forest litter, Alnus   , ferns, Verathrum, 12.vii.1998 (V.I.Gusarov) ( SPSU) GoogleMaps   .

Diagnosis. See Klimaszewski and Winchester (2002; as Atheta vasta   ).

Discussion. The lectotype of At. innocens   agrees with the types of At. keeni   in external characters and the shape of the spermatheca. The lectotype of At. profecta   has no head and prothorax but agrees with the types of At. keeni   in the shape of the aedeagus and abdominal tergum 8. The lectotype of At. profecta   is poorly pigmented, but otherwise it agrees completely with At. keeni   in external characters, including male secondary features, and in the shape of the aedeagus.

The type series of Homalota vasta Mäklin   in Mannerheim, 1853 (examined) includes two specimens ( Lohse & Smetana 1985). One of them is a female conspecific with Atheta keeni   and the other is a female belonging to the genus Liogluta   and illustrated by Lohse and Smetana (1985). This second female bears the following labels: “Sitka”, “Holmberg”, “Mus. Zool. H:fors. Spec. Typ. No. 2251. Homalota vasta Mäklin   ”. From the description and illustrations in Lohse and Smetana (1985) it is clear that they meant to designate the second specimen (a female of Liogluta   ) as the lectotype of Homalota vasta   . Unfortunately, when they cited the labels of the lectotype, they listed the labels of the first specimen (a female of Atheta keeni   ). Moreover, it is the first specimen that had been labeled as the lectotype. The latter circumstance is especially misleading and originally I was going to accept what the lectotype label said, and consider Atheta keeni   and Homalota vasta   to be synonymous. While identifying some athetines from British Columbia for Jan Klimaszewski ( Klimaszewski & Winchester 2002) I labeled the specimens of Atheta keeni   as At. vasta   . However, Lohse and Smetana are very explicit in that they meant the specimen of Liogluta   , illustrated and redescribed by them, to be the lectotype of Homalota vasta   . Therefore I accept their intent to designate the Liogluta   specimen as the lectotype, disregard their misplaced lectotype label and consider At. keeni   to be a valid species, different from Liogluta vasta   . In the type series of Homalota vasta   I labeled the specimen of Liogluta   as lectotype and the specimen of At. keeni   as paralectotype.

Atheta keeni   is the type species of the subgenus Lamiota Casey, 1910a   (by original designation). Subgeneric assignment of At. keeni   and the status of the name Lamiota   requires further study. Atheta keeni   has the pronotal pubescence of the type I and cannot be placed in the genus Liogluta Thomson, 1858   .

Distribution. Atheta keeni   is known from Alaska, British Columbia and Oregon.

NMNH

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Atheta

Loc

Atheta keeni Casey, 1910a

Gusarov, Vladimir I. 2003
2003
Loc

Atheta (Liogluta) vasta: Klimaszewski & Winchester, 2002: 39

Klimaszewski, J. & Winchester, N. N. & K. Bolte 2002: 39
Lohse, G. A. & Smetana, A. 1985: 297
2002
Loc

Lamiota keeni:

Seevers, C. H. 1978: 263
1978
Loc

Xenota innocens:

Seevers, C. H. 1978: 270
1978
Loc

Lamiota achromata:

Seevers, C. H. 1978: 263
1978
Loc

Lamiota profecta:

Seevers, C. H. 1978: 263
1978
Loc

Atheta (Liogluta) keeni: Moore & Legner, 1975: 364

Moore, I. & Legner, E. F. 1975: 364
1975
Loc

Atheta (s. str.) innocens: Moore & Legner, 1975: 363

Moore, I. & Legner, E. F. 1975: 363
1975
Loc

Atheta (Liogluta) achromata: Moore & Legner, 1975: 353

Moore, I. & Legner, E. F. 1975: 353
1975
Loc

Atheta (Liogluta) profecta: Moore & Legner, 1975: 370

Moore, I. & Legner, E. F. 1975: 370
1975
Loc

Atheta (Liogluta) keeni:

Bernhauer, M. & Scheerpeltz, O. 1926: 657
1926
Loc

Atheta (s. str.) innocens:

Bernhauer, M. & Scheerpeltz, O. 1926: 644
1926
Loc

Atheta (Liogluta) achromata:

Bernhauer, M. & Scheerpeltz, O. 1926: 656
1926
Loc

Atheta (Liogluta) profecta:

Bernhauer, M. & Scheerpeltz, O. 1926: 658
1926
Loc

Atheta (Liogluta) keeni:

Fenyes, A. 1920: 209
1920
Loc

Atheta (s. str.) innocens:

Fenyes, A. 1920: 214
1920
Loc

Atheta (Liogluta) achromata:

Fenyes, A. 1920: 209
1920
Loc

Atheta (Liogluta) profecta:

Fenyes, A. 1920: 209
1920
Loc

Atheta (Lamiota) achromata

Casey, T. L. 1911: 82
1911
Loc

Atheta (Lamiota) profecta

Casey, T. L. 1911: 83
1911
Loc

Atheta (Lamiota) keeni

Casey, T. L. 1910: 17
1910
Loc

Atheta (s. str.) innocens

Casey, T. L. 1910: 39
1910