Atheta (Dimetrota auct.) picipennis ( Mannerheim, 1843 )

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134: 40-42

publication ID

http://doi.org/ 10.11646/zootaxa.353.1.1

publication LSID

lsid:zoobank.org:pub:104E9C0B-B207-41E6-816C-E91FC1C7DBA0

DOI

http://doi.org/10.5281/zenodo.5102684

persistent identifier

http://treatment.plazi.org/id/03F387B2-FFA7-5365-CD24-50E4FEA1A721

treatment provided by

Felipe

scientific name

Atheta (Dimetrota auct.) picipennis ( Mannerheim, 1843 )
status

 

Atheta (Dimetrota auct.) picipennis ( Mannerheim, 1843)  

( Figs. 3 View FIGURES 1–4 , 32 View FIGURES 31–35 in Lohse & Smetana 1985; Figs. 63–66 View FIGURES 60–63 View FIGURES 64–69 in Klimaszewski & Winchester 2002)

Homalota picipennis Mannerheim, 1843: 224   .

Atheta (s. str.) carlottae Casey, 1910a: 32   .

Atheta (s. str.) aperta Casey, 1910a: 33   , syn. nov.

Atheta (s. str.) wrangelica Casey, 1911a: 91   , syn. nov.

Atheta (s. str.) morbosa Casey, 1911a: 107   , syn. nov.

Atheta (s. str.) intacta Casey, 1911a: 110   , syn. nov.

Atheta (s. str.) alaskana Casey, 1911a: 113   , syn. nov.

Atheta (Dimetrota) picipennis: Fenyes, 1920: 204   (ex parte; as synonym of At. rugulosa ( Heer, 1839))   .

Atheta (s. str.) carlottae: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) aperta: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) wrangelica: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) morbosa: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) intacta: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) alaskana: Fenyes, 1920: 215   (as valid species).

Atheta (Dimetrota) picipennis: Bernhauer & Scheerpeltz, 1926: 664   (ex parte; as valid species).

Atheta (s. str.) carlottae: Bernhauer & Scheerpeltz, 1926: 639   (as valid species).

Atheta (s. str.) aperta: Bernhauer & Scheerpeltz, 1926: 638   (as valid species).

Atheta (s. str.) wrangelica: Bernhauer & Scheerpeltz, 1926: 652   (as valid species).

Atheta (s. str.) morbosa: Bernhauer & Scheerpeltz, 1926: 645   (as valid species).

Atheta (s. str.) intacta: Bernhauer & Scheerpeltz, 1926: 644   (as valid species).

Atheta (s. str.) alaskana: Bernhauer & Scheerpeltz, 1926: 638   (as valid species).

Atheta (Dimetrota) picipennis: Moore & Legner, 1975: 370   (ex parte; as valid species).

Atheta (s. str.) carlottae: Moore & Legner, 1975: 356   (as valid species).

Atheta (s. str.) aperta: Moore & Legner, 1975: 354   (as valid species; misspelled as apera).

Atheta (s. str.) wrangelica: Moore & Legner, 1975: 377   (as valid species).

Atheta (s. str.) morbosa: Moore & Legner, 1975: 366   (as valid species).

Atheta (s. str.) intacta: Moore & Legner, 1975: 364   (as valid species).

Atheta (s. str.) alaskana: Moore & Legner, 1975: 354   (as valid species).

Dimetrota picipennis: Seevers, 1978: 260   (as valid species).

Xenota carlottae: Seevers, 1978: 270   (as valid species).

Xenota aperta: Seevers, 1978: 269   (as valid species).

Xenota wrangelica: Seevers, 1978: 271   (as valid species).

Xenota morbosa: Seevers, 1978: 270   (as valid species).

Xenota intacta: Seevers, 1978: 270   (as valid species).

Xenota alaskana: Seevers, 1978: 269   (as valid species).

Dimetrota picipennis: Lohse & Smetana, 1985: 283   (as valid species).

Dimetrota carlottae: Lohse & Smetana, 1985: 283   (as synonym of Dim. picipennis   ).

Atheta (Dimetrota) picipennis: Klimaszewski & Winchester, 2002: 26   (as valid species).

Type material. Lectotype of Homalota picipennis   (designated by Lohse and Smetana (1985)): UNITED STATES: Alaska: ♂, Sitka ( MZHF).  

Lectotype of Atheta carlottae   (here designated): ♂, “QCI [ Queen Charlotte Islands ]”, “ Atheta carlottae Csy.   ”, “TYPE USNM 39247” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: 2♂♂, 2♀♀, “QCI [ Queen Charlotte Islands ]”, “ carlottae   PARATYPE USNM 39247” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta aperta   (here designated): ♂, “Cal.[ifornia, Humboldt Co., Arcata ]”, “ Atheta aperta Csy.   ”, “TYPE USNM 39250” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotype: ♀, “Cal.[ifornia, Humboldt Co., Arcata ]”, “ aperta   ­2 PARATYPE USNM 39250” (red label), “CASEY bequest 1925” ( NMNH)   .

Holotype of Atheta wrangelica   : ♀, “ Ft. Wrangel [Wrangell], Alaska. Wickham”, “ ♀ ”, “ wrangelica Csy.   ”, “TYPE USNM 39248” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta morbosa   (here designated): ♀, “Cal.[ifornia, Saucelito near San Francisco]”, “ Atheta morbosa Csy.   ”, “TYPE USNM 39293” (red label), “CASEY bequest 1925” ( NMNH).  

Holotype of Atheta intacta   : ♀, “Cal.[ifornia, Marin Co., Saucelito ]”, “ intacta Csy.   ”, “TYPE USNM 39299” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta alaskana   (here designated): ♀, “Alask.[Alaska, “without further indication, but probably collected on the mainland near Fort Wrangel” ( Casey 1910a)] Wick.[ham]”, “ alaskana Csy.   ”, “TYPE USNM 39281” (red label), “CASEY bequest 1925” ( NMNH).

Additional material: CANADA: British Columbia: ♂, Upper Carmanah Valley , 31.vii–11.viii.1991 (N.Winchester)   ; ♀, ditto but 28.viii–9.ix.1991 ( SPSU)   ; UNITED STATES: California: San Francisco Co.: ♀ ( NMNH (Casey collection))   .

Diagnosis. See Lohse & Smetana (1985); Klimaszewski & Winchester (2002).

Discussion. The name Atheta picipennis ( Mannerheim, 1843)   (ex Homalota   ) is a secondary junior homonym of the name Atheta picipennis ( Stephens, 1832)   (ex Aleochara   ) which is itself a primary junior homonym of the name Aleochara picipennis Gyllenhal, 1827   . Atheta picipennis ( Stephens, 1832)   has been considered a synonym of Atheta amicula ( Stephens, 1832)   since at least 1868 ( Gemminger & Harold 1868) and to the best of my knowledge has not been cited as valid after 1899 (Article 23.9.1.1). In the last 50 years at least 31 works by at least 42 authors have been published ( Hatch 1957; Gensicke 1960; Hansen et. al. 1960; Strand & Vik 1964; Koch 1968; Scheerpeltz 1968; Easton 1970; Palm 1970; Blackwelder 1973; Bordoni 1973; Tikhomirova 1973; Benick & Lohse 1974; Moore & Legner 1975; Yosii & Sawada 1976; Hanski & Koskela 1977; Pope 1977; Hanski & Koskela 1978; Muona 1979b; Burakowski et al. 1981; Uhlig & Vogel 1981; Benick 1982; Sawada 1982; Muona 1984; Lohse 1985; Segers 1986; Kahlen 1987; Muona 1987; Silfverberg 1988; Alexandrovitch et al. 1996; Derunkov & Melke 2001; Klimaszewski & Winchester 2002) listing Atheta picipennis   as valid (Article 23.9.1.2). Atheta picipennis ( Mannerheim, 1843)   is a nomen protectum and Atheta picipennis ( Stephens, 1832)   is a nomen oblitum (Article 23.9.2).

Atheta picipennis   was believed to have a Holarctic distribution (e.g., Bernhauer & Scheerpeltz 1926) until Lohse and Smetana (1985) demonstrated that the Palaearctic specimens belonged to a separate species, At. aeneipennis ( Thomson, 1856)   .

Females of At. picipennis   and At. altaica Bernhauer, 1901   have very similar spermathecae. Klimaszewski and Winchester (2002) were unable to find any difference between females of the two species. Although in average At. picipennis   has a larger body than At. altaica   , females of these two species can be reliably identified only by association with males. In comparison to At. altaica   , males of At. picipennis   have a narrower apex of the median lobe of the aedeagus.

Distribution. Atheta picipennis   and At. altaica   appear to be allopatric: Atheta picipennis   is known from southeast Alaska (Sitka, Wrangell), coastal British Columbia and California, while At. altaica   is known (in North America) from the Kenai Peninsula (Alaska), central Alaska, Yukon Territory, Northwest Territories and Labrador ( Lohse et al. 1990).

NMNH

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Atheta

Loc

Atheta (Dimetrota auct.) picipennis ( Mannerheim, 1843 )

Gusarov, Vladimir I. 2003
2003
Loc

Atheta (Dimetrota) picipennis:

Klimaszewski, J. & Winchester, N. N. & K. Bolte 2002: 26
2002
Loc

Dimetrota picipennis: Lohse & Smetana, 1985: 283

Lohse, G. A. & Smetana, A. 1985: 283
1985
Loc

Dimetrota carlottae:

Lohse, G. A. & Smetana, A. 1985: 283
1985
Loc

Dimetrota picipennis:

Seevers, C. H. 1978: 260
1978
Loc

Xenota carlottae:

Seevers, C. H. 1978: 270
1978
Loc

Xenota aperta:

Seevers, C. H. 1978: 269
1978
Loc

Xenota wrangelica:

Seevers, C. H. 1978: 271
1978
Loc

Xenota morbosa:

Seevers, C. H. 1978: 270
1978
Loc

Xenota intacta:

Seevers, C. H. 1978: 270
1978
Loc

Xenota alaskana:

Seevers, C. H. 1978: 269
1978
Loc

Atheta (Dimetrota) picipennis:

Moore, I. & Legner, E. F. 1975: 370
1975
Loc

Atheta (s. str.) carlottae: Moore & Legner, 1975: 356

Moore, I. & Legner, E. F. 1975: 356
1975
Loc

Atheta (s. str.) aperta: Moore & Legner, 1975: 354

Moore, I. & Legner, E. F. 1975: 354
1975
Loc

Atheta (s. str.) wrangelica: Moore & Legner, 1975: 377

Moore, I. & Legner, E. F. 1975: 377
1975
Loc

Atheta (s. str.) morbosa: Moore & Legner, 1975: 366

Moore, I. & Legner, E. F. 1975: 366
1975
Loc

Atheta (s. str.) intacta: Moore & Legner, 1975: 364

Moore, I. & Legner, E. F. 1975: 364
1975
Loc

Atheta (s. str.) alaskana: Moore & Legner, 1975: 354

Moore, I. & Legner, E. F. 1975: 354
1975
Loc

Atheta (Dimetrota) picipennis:

Bernhauer, M. & Scheerpeltz, O. 1926: 664
1926
Loc

Atheta (s. str.) carlottae:

Bernhauer, M. & Scheerpeltz, O. 1926: 639
1926
Loc

Atheta (s. str.) aperta:

Bernhauer, M. & Scheerpeltz, O. 1926: 638
1926
Loc

Atheta (s. str.) wrangelica:

Bernhauer, M. & Scheerpeltz, O. 1926: 652
1926
Loc

Atheta (s. str.) morbosa:

Bernhauer, M. & Scheerpeltz, O. 1926: 645
1926
Loc

Atheta (s. str.) intacta:

Bernhauer, M. & Scheerpeltz, O. 1926: 644
1926
Loc

Atheta (s. str.) alaskana:

Bernhauer, M. & Scheerpeltz, O. 1926: 638
1926
Loc

Atheta (Dimetrota) picipennis:

Fenyes, A. 1920: 204
1920
Loc

Atheta (s. str.) carlottae:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) aperta:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) wrangelica:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) morbosa:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) intacta:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) alaskana:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) wrangelica

Casey, T. L. 1911: 91
1911
Loc

Atheta (s. str.) morbosa

Casey, T. L. 1911: 107
1911
Loc

Atheta (s. str.) intacta

Casey, T. L. 1911: 110
1911
Loc

Atheta (s. str.) alaskana

Casey, T. L. 1911: 113
1911
Loc

Atheta (s. str.) carlottae

Casey, T. L. 1910: 32
1910
Loc

Atheta (s. str.) aperta

Casey, T. L. 1910: 33
1910
Loc

Homalota picipennis

Mannerheim, C. G. 1843: 224
1843