Atheta (crassicornis group) modesta ( Melsheimer, 1844 )

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134: 50-54

publication ID

http://doi.org/ 10.11646/zootaxa.353.1.1

publication LSID

lsid:zoobank.org:pub:104E9C0B-B207-41E6-816C-E91FC1C7DBA0

DOI

http://doi.org/10.5281/zenodo.5102694

persistent identifier

http://treatment.plazi.org/id/03F387B2-FFB1-5379-CD24-524EFB7BA4E7

treatment provided by

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scientific name

Atheta (crassicornis group) modesta ( Melsheimer, 1844 )
status

 

Atheta (crassicornis group) modesta ( Melsheimer, 1844)  

( Figs. 56–69 View FIGURES 56–59 View FIGURES 60–63 View FIGURES 64–69 )

Homalota modesta Melsheimer, 1844: 31   .

Atheta (Acrotona) modesta: Bernhauer, 1907: 398   (as valid species).

Atheta (s. str.) crassicornis var. virginica Bernhauer, 1907: 393   , syn. nov.

Atheta virginica: Blatchley, 1910: 356   (as valid species).

Atheta (s. str.) rhodeana Casey, 1910a: 24   , syn. nov.

Atheta (s. str.) capella Casey, 1910a: 24   , syn. nov.

Atheta (s. str.) ducens Casey, 1910a: 25   , syn. nov.

Atheta (s. str.) temperans Casey, 1910a: 27   , syn. nov.

Atheta (Delphota) logica Casey, 1911a: 86   , syn. nov.

Atheta (s. str.) tradita Casey, 1911a: 101   , syn. nov.

Atheta (s. str.) fenisex Casey, 1911a: 102   , syn. nov.

Atheta (s. str.) vierecki Casey, 1911a: 103   , syn. nov.

Atheta (s. str.) auguralis Casey, 1911a: 104   , syn. nov.

Atheta (s. str.) bifaria Casey, 1911a: 104   , syn. nov.

Atheta (Acrotona) modesta: Fenyes, 1920: 225   (as valid species).

Atheta (s. str.) virginica: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) rhodeana: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) capella: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) ducens: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) temperans: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) logica: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) tradita: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) fenisex: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) vierecki: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) auguralis: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) bifaria: Fenyes, 1920: 215   (as valid species).

Atheta (Acrotona) modesta: Bernhauer & Scheerpeltz, 1926: 675   (as valid species).

Atheta (s. str.) virginica: Bernhauer & Scheerpeltz, 1926: 652   (as valid species).

Atheta (s. str.) rhodeana: Bernhauer & Scheerpeltz, 1926: 649   (as valid species).

Atheta (s. str.) capella: Bernhauer & Scheerpeltz, 1926: 639   (as valid species).

Atheta (s. str.) ducens: Bernhauer & Scheerpeltz, 1926: 642   (as valid species).

Atheta (s. str.) temperans: Bernhauer & Scheerpeltz, 1926: 650   (as valid species).

Atheta (s. str.) logica: Bernhauer & Scheerpeltz, 1926: 645   (as valid species).

Atheta (s. str.) tradita: Bernhauer & Scheerpeltz, 1926: 651   (as valid species).

Atheta (s. str.) fenisex: Bernhauer & Scheerpeltz, 1926: 642   (as valid species).

Atheta (s. str.) vierecki: Bernhauer & Scheerpeltz, 1926: 651   (as valid species).

Atheta (s. str.) auguralis: Bernhauer & Scheerpeltz, 1926: 638   (as valid species).

Atheta (s. str.) bifaria: Bernhauer & Scheerpeltz, 1926: 639   (as valid species).

Atheta (Acrotona) modesta: Moore & Legner, 1975: 366   (as valid species).

Atheta (s. str.) virginica: Moore & Legner, 1975: 377   (as valid species).

Atheta (s. str.) rhodeana: Moore & Legner, 1975: 372   (as valid species).

Atheta (s. str.) capella: Moore & Legner, 1975: 356   (as valid species).

Atheta (s. str.) ducens: Moore & Legner, 1975: 359   (as valid species).

Atheta (s. str.) temperans: Moore & Legner, 1975: 366   (as valid species; misspelled as temporans). Atheta (s. str.) logica: Moore & Legner, 1975: 365   (as valid species).

Atheta (s. str.) tradita: Moore & Legner, 1975: 376   (as valid species).

Atheta (s. str.) fenisex: Moore & Legner, 1975: 360   (as valid species).

Atheta (s. str.) vierecki: Moore & Legner, 1975: 377   (as valid species).

Atheta (s. str.) auguralis: Moore & Legner, 1975: 354   (as valid species; misspelled as angularis   ). Atheta (s. str.) bifaria: Moore & Legner, 1975: 355   (as valid species).

Acrotona (s. str.) modesta: Seevers, 1978: 256   (as valid species).

Xenota virginica: Seevers, 1978: 269   (as valid species).

Xenota rhodeana: Seevers, 1978: 269   (as valid species).

Xenota capella: Seevers, 1978: 268   (as valid species).

Xenota ducens: Seevers, 1978: 268   (as valid species).

Xenota temperans: Seevers, 1978: 269   (as valid species).

Xenota logica: Seevers, 1978: 268   (as valid species).

Xenota tradita: Seevers, 1978: 269   (as valid species).

Xenota fenisex: Seevers, 1978: 268   (as valid species).

Xenota vierecki: Seevers, 1978: 269   (as valid species).

Xenota auguralis: Seevers, 1978: 267   (as valid species).

Xenota bifaria: Seevers, 1978: 267   (as valid species).

Type material. Lectotype of Homalota modesta   (here designated): ♂, “U.[nited] S.[tates]”, “ modesta Melsh.   ”, “Melsh.[eimer]”, “ modesta   ”, a red piece of paper, “MCZ TYPE 34936” (red label) ( MCZ) (Two paralectotypes of H. modesta   (with labels “ modesta   ”, “Melsh.”, “ MCZ TYPE 34936”) are not conspecific with the lectotype and belong to Stethusa spuriella ( Casey, 1910a) (MCZ))   .

Lectotype of Atheta crassicornis var. virginica   (here designated): ♂, “White Sulphur W.[est] Va. [Virginia]”, “Aug.[ust]”, yellow circle, “ pacifica Bh. Fenyes det. Bernhauer”, “ v. virginica Brh.   Typus” (yellow label), “Chicago NHMus M.Bernhauer Collection” ( FMNH) (The paralectotype of this species (♂, “E. Machias, Me.[Maine]”, “June”, white circle, “ crassicornis Gvh. Fenyes   det. Bernhauer”, “ v. virginica Brh. Cotypus   ” (yellow label), “Chicago NHMus M.Bernhauer Collection” ( FMNH)) is not conspecific with the lectotype. It belongs to At. crenuliventris Bernhauer, 1907   ).

Lectotype of Atheta rhodeana   (here designated): ♂, “R.[hode] I.[sland, Boston Neck]”, “ rhodeana   ­2 PARATYPE USNM 39215” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: ♂, “R.[hode] I.[sland, Boston Neck]”, “ ♂ ”, “ Atheta rhodeana Csy.   ”, “TYPE USNM 39215” (red label), “CASEY bequest 1925” ( NMNH)   ; 6♀♀, “R.[hode] I.[sland, Boston Neck]”, “ rhodeana   PARATYPE USNM 39215” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta capella   (here designated): ♂, “R.[hode] I.[sland, Boston Neck]”, “ capella   ­2 PARATYPE USNM 39216” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: ♂ (with the aedeagus missing), “R.[hode] I.[sland, Boston Neck]”, “ Atheta capella Csy.   ”, “TYPE USNM 39216” (red label), “CASEY bequest 1925” ( NMNH)   ; 2♂♂, ♀, “R.[hode] I.[sland, Boston Neck]”, “ capella   PARATYPE USNM 39216” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta ducens   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ ducens Csy.   ”, “TYPE USNM 39218” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: 17 specimens, “N.[ew] Y.[ork, Catskill Mts. ]”, “ ducens   PARATYPE USNM 39218” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta temperans   (here designated): ♀, “D.[istrict of] C.[olumbia]”, “ Atheta temperans Csy.   ”, “TYPE USNM 39225” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta logica   (here designated): ♀, “Penn.[sylvania, near Philadelphia]”, “ logica Csy.   ”, “TYPE USNM 39489” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotype: ♀, “ Penn.[sylvania, near Philadelphia]”, “ logica   PARATYPE USNM 39489” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta tradita   (here designated): ♀, “N.[ew] Y.[ork, Catskill Mts. ]”, “ tradita Csy.   ”, “TYPE USNM 39253” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotype: ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ tradita   PARATYPE USNM 39253” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta fenisex   (here designated): ♀, “R.[hode] I.[sland, Boston Neck]”, “ fenisex Csy.   ”, “TYPE USNM 39256” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta vierecki   (here designated): ♂, “Double Beach, Ct. [Connecticut] 19 Aug. 1905 H.L.Viereck ”, “Fungus”, “ vierecki Csy.   ”, “TYPE USNM 39255” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: 2♂♂, ♀, “Double Beach, Ct. [Connecticut] 19 Aug. 1905 H.L.Viereck ”, “Fungus”, “ vierecki   PARATYPE USNM 39255” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta auguralis   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ auguralis Csy.   ”, “TYPE USNM 39257” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta bifaria   (here designated): ♀, “D.[istrict of] C.[olumbia]”, “ bifaria Csy.   ”, “TYPE USNM 39258” (red label), “CASEY bequest 1925” ( NMNH).  

Additional material. UNITED STATES: Michigan: Newaygo Co.: ♂, 2♀♀, 16 km SEE White Cloud, Newaygo State Park, 43º29.84'N 85º34.85'W, 150 m, in forest litter, Pinus   , Quercus   , 16.vii.1999 (V.I.Gusarov) ( SPSU); New York: Orange Co.: ♀, 8 km S New Windsor, Black Rock Forest, 9.v.1998 (V.I.Gusarov); ♀, ditto but 23–25.v.1998; ♂, ditto but pitfall traps, vi.1998 (V.Ovcharenko) ( SPSU); Sullivan Co.: ♀, Jeffersonville, 11–12.ix.1999 (A.Berkov) ( SPSU); Pennsylvania: 3♂♂, 4♀♀, “Penn.[sylvania, near Philadelphia]”, ( NMNH (Casey collection)); Vermont: Windsor Co.: 2♀♀, 8 km SEE Springfield, Hwy. I­91, 43º14.00'N 72º26.81'W, 250 m, in forest litter, Acer   , Betula   , Tsuga   , Quercus   , 21.ix.1998 (V.I.Gusarov) ( SPSU). CANADA: New Brunswick: Sunbury Co.: ♂, ♀, Acadia Research Forest, pitfall traps, 4.viii.1999 (G.Gesner); ♀, ditto but 13.vii.1999 ( SPSU); Ontario: ♂, ♀, 22 km NNE Gananoque, env. of Charleston Lake Prov. Park, 44º30.03'N 76º03.96'W, 0 m, in forest litter, Acer   , Ulmus   , Quercus   , Pinus   , ferns, 15.vii.1999 (V.I.Gusarov) ( SPSU); Québec: ♂, Parc de la Gatineau, Lac Lapêche, 9 km SWW Sainte­François­de­Masham, 26.ix.1998 (V.I.Gusarov); ♂, St.­Jaques­de­Leeds, 46º16'N 71º23'W, maple forest, 2.viii.1993 ( SPSU); 5 specimens, ditto but 15.vii.1994 ( LFC); ♀, ditto but 15.vii.1994; ♀, ditto but 19.vii.1994; ♂, 170 km NE Québec­City, Pohénégamook, 47º28'N 69º14'W, maple forest, 19.vii.1994 ( SPSU).

Diagnosis. Atheta modesta   is closely related to the Palaearctic At. crassicornis ( Fabricius, 1793)   but differs in having the pronotal setation pattern of the type III, the distinct shape of the male tergum 8 ( Fig. 56 View FIGURES 56–59 ) and in the shape of the aedeagus ( Figs. 60– 68 View FIGURES 60–63 View FIGURES 64–69 ). The spermathecae of the two species are very similar ( Fig. 69 View FIGURES 64–69 ).

Discussion. Bernhauer (1907) described At. virginica   as a variety of At. crassicornis   . He noted the similarity between the examined North American specimens and the European specimens, but listed several differences between the two. He also noted that when additional Nearctic specimens become available they might prove to represent a distinct species and not a variety of At. crassicornis   .

The types of H. modesta   , At. crassicornis var. virginica   , At. rhodeana   , At. capella   , At. ducens   , At. temperans   , At. logica   , At. tradita   , At. fenisex   , At. vierecki   , At. auguralis   and At. bifaria   are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.

Distribution. Atheta modesta   is widely distributed in the eastern United States (Michigan, Vermont, Rhode Island, Connecticut, New York, Pennsylvania, West Virginia and District of Columbia) and Canada (New Brunswick, Ontario and Québec).

MCZ

Museum of Comparative Zoology

FMNH

Field Museum of Natural History

NMNH

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Atheta

Loc

Atheta (crassicornis group) modesta ( Melsheimer, 1844 )

Gusarov, Vladimir I. 2003
2003
Loc

Acrotona (s. str.) modesta:

Seevers, C. H. 1978: 256
1978
Loc

Xenota virginica: Seevers, 1978: 269

Seevers, C. H. 1978: 269
1978
Loc

Xenota rhodeana:

Seevers, C. H. 1978: 269
1978
Loc

Xenota capella:

Seevers, C. H. 1978: 268
1978
Loc

Xenota ducens:

Seevers, C. H. 1978: 268
1978
Loc

Xenota temperans:

Seevers, C. H. 1978: 269
1978
Loc

Xenota logica:

Seevers, C. H. 1978: 268
1978
Loc

Xenota tradita:

Seevers, C. H. 1978: 269
1978
Loc

Xenota fenisex:

Seevers, C. H. 1978: 268
1978
Loc

Xenota vierecki:

Seevers, C. H. 1978: 269
1978
Loc

Xenota auguralis:

Seevers, C. H. 1978: 267
1978
Loc

Xenota bifaria:

Seevers, C. H. 1978: 267
1978
Loc

Atheta (Acrotona) modesta:

Moore, I. & Legner, E. F. 1975: 366
1975
Loc

Atheta (s. str.) virginica:

Moore, I. & Legner, E. F. 1975: 377
1975
Loc

Atheta (s. str.) rhodeana: Moore & Legner, 1975: 372

Moore, I. & Legner, E. F. 1975: 372
1975
Loc

Atheta (s. str.) capella: Moore & Legner, 1975: 356

Moore, I. & Legner, E. F. 1975: 356
1975
Loc

Atheta (s. str.) ducens: Moore & Legner, 1975: 359

Moore, I. & Legner, E. F. 1975: 359
1975
Loc

Atheta (s. str.) temperans: Moore & Legner, 1975: 366

Moore, I. & Legner, E. F. 1975: 366
Moore, I. & Legner, E. F. 1975: 365
1975
Loc

Atheta (s. str.) tradita: Moore & Legner, 1975: 376

Moore, I. & Legner, E. F. 1975: 376
1975
Loc

Atheta (s. str.) fenisex: Moore & Legner, 1975: 360

Moore, I. & Legner, E. F. 1975: 360
1975
Loc

Atheta (s. str.) vierecki: Moore & Legner, 1975: 377

Moore, I. & Legner, E. F. 1975: 377
1975
Loc

Atheta (s. str.) auguralis: Moore & Legner, 1975: 354

Moore, I. & Legner, E. F. 1975: 354
Moore, I. & Legner, E. F. 1975: 355
1975
Loc

Atheta (Acrotona) modesta:

Bernhauer, M. & Scheerpeltz, O. 1926: 675
1926
Loc

Atheta (s. str.) virginica:

Bernhauer, M. & Scheerpeltz, O. 1926: 652
1926
Loc

Atheta (s. str.) rhodeana:

Bernhauer, M. & Scheerpeltz, O. 1926: 649
1926
Loc

Atheta (s. str.) capella:

Bernhauer, M. & Scheerpeltz, O. 1926: 639
1926
Loc

Atheta (s. str.) ducens:

Bernhauer, M. & Scheerpeltz, O. 1926: 642
1926
Loc

Atheta (s. str.) temperans:

Bernhauer, M. & Scheerpeltz, O. 1926: 650
1926
Loc

Atheta (s. str.) logica:

Bernhauer, M. & Scheerpeltz, O. 1926: 645
1926
Loc

Atheta (s. str.) tradita:

Bernhauer, M. & Scheerpeltz, O. 1926: 651
1926
Loc

Atheta (s. str.) fenisex:

Bernhauer, M. & Scheerpeltz, O. 1926: 642
1926
Loc

Atheta (s. str.) vierecki:

Bernhauer, M. & Scheerpeltz, O. 1926: 651
1926
Loc

Atheta (s. str.) auguralis:

Bernhauer, M. & Scheerpeltz, O. 1926: 638
1926
Loc

Atheta (s. str.) bifaria:

Bernhauer, M. & Scheerpeltz, O. 1926: 639
1926
Loc

Atheta (Acrotona) modesta:

Fenyes, A. 1920: 225
1920
Loc

Atheta (s. str.) virginica:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) rhodeana:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) capella:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) ducens:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) temperans:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) logica:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) tradita:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) fenisex:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) vierecki:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) auguralis:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) bifaria:

Fenyes, A. 1920: 215
1920
Loc

Atheta (Delphota) logica

Casey, T. L. 1911: 86
1911
Loc

Atheta (s. str.) tradita

Casey, T. L. 1911: 101
1911
Loc

Atheta (s. str.) fenisex

Casey, T. L. 1911: 102
1911
Loc

Atheta (s. str.) vierecki

Casey, T. L. 1911: 103
1911
Loc

Atheta (s. str.) auguralis

Casey, T. L. 1911: 104
1911
Loc

Atheta (s. str.) bifaria

Casey, T. L. 1911: 104
1911
Loc

Atheta virginica:

Blatchley, W. S. 1910: 356
1910
Loc

Atheta (s. str.) rhodeana

Casey, T. L. 1910: 24
1910
Loc

Atheta (s. str.) capella

Casey, T. L. 1910: 24
1910
Loc

Atheta (s. str.) ducens

Casey, T. L. 1910: 25
1910
Loc

Atheta (s. str.) temperans

Casey, T. L. 1910: 27
1910
Loc

Atheta (Acrotona) modesta:

Bernhauer, M. 1907: 398
1907
Loc

Atheta (s. str.) crassicornis var. virginica

Bernhauer, M. 1907: 393
1907
Loc

Homalota modesta

Melsheimer, F. E. 1844: 31
1844