Atheta (Tetropla) frosti Bernhauer, 1909

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134: 55-60

publication ID

http://doi.org/ 10.11646/zootaxa.353.1.1

publication LSID

lsid:zoobank.org:pub:104E9C0B-B207-41E6-816C-E91FC1C7DBA0

DOI

http://doi.org/10.5281/zenodo.5103050

persistent identifier

http://treatment.plazi.org/id/03F387B2-FFB6-5377-CD24-5517FCBBA0A1

treatment provided by

Felipe

scientific name

Atheta (Tetropla) frosti Bernhauer, 1909
status

 

Atheta (Tetropla) frosti Bernhauer, 1909  

( Figs. 70–80 View FIGURES 70–74 View FIGURES 75–80 )

Atheta (s. str.) nigritula: Bernhauer, 1907: 392   (misidentification).

Atheta (s. str.) frosti Bernhauer, 1909: 520   .

Atheta (s. str.) comitata Casey, 1910a: 24   , syn. nov.

Atheta (s. str.) gnoma Casey, 1910a: 25   , syn. nov.

Atheta (s. str.) elota Casey, 1910a: 26   , syn. nov.

Atheta (s. str.) insidiosa Casey, 1910a: 26   , syn. nov.

Pseudota puricula Casey, 1911a: 148   , syn. nov.

Atheta (s. str.) candidula Casey, 1911a: 94   , syn. nov.

Atheta (s. str.) diffisa Casey, 1911a: 95   , syn. nov.

Atheta (s. str.) nata Casey, 1911a: 95   , syn. nov.

Atheta (s. str.) modiella Casey, 1911a: 96   , syn. nov.

Atheta (s. str.) vacillans Casey, 1911a: 98   , syn. nov.

Atheta (s. str.) frosti: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) comitata: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) gnoma: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) elota: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) insidiosa: Fenyes, 1920: 213   (as valid species).

Atheta (Pancota) puricula: Fenyes, 1920: 202   (as valid species).

Atheta (s. str.) candidula: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) diffisa: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) nata: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) modiella: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) vacillans: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) frosti: Bernhauer & Scheerpeltz, 1926: 643   (as valid species).

Atheta (s. str.) comitata: Bernhauer & Scheerpeltz, 1926: 640   (as valid species).

Atheta (s. str.) gnoma: Bernhauer & Scheerpeltz, 1926: 643   (as valid species).

Atheta (s. str.) elota: Bernhauer & Scheerpeltz, 1926: 642   (as valid species).

Atheta (s. str.) insidiosa: Bernhauer & Scheerpeltz, 1926: 644   (as valid species).

Atheta (Pancota) puricula: Bernhauer & Scheerpeltz, 1926: 661   (as valid species).

Atheta (s. str.) candidula: Bernhauer & Scheerpeltz, 1926: 639   (as valid species).

Atheta (s. str.) diffisa: Bernhauer & Scheerpeltz, 1926: 641   (as valid species).

Atheta (s. str.) nata: Bernhauer & Scheerpeltz, 1926: 646   (as valid species).

Atheta (s. str.) modiella: Bernhauer & Scheerpeltz, 1926: 645   (as valid species).

Atheta (s. str.) vacillans: Bernhauer & Scheerpeltz, 1926: 651   (as valid species).

Atheta (s. str.) frosti: Moore & Legner, 1975: 361   (as valid species).

Atheta (s. str.) comitata: Moore & Legner, 1975: 357   (as valid species).

Atheta (s. str.) gnoma: Moore & Legner, 1975: 362   (as valid species).

Atheta (s. str.) elota: Moore & Legner, 1975: 360   (as valid species).

Atheta (s. str.) insidiosa: Moore & Legner, 1975: 363   (as valid species).

Atheta (Pancota) puricula: Moore & Legner, 1975: 371   (as valid species; misspelled as furicula). Atheta (s. str.) candidula: Moore & Legner, 1975: 356   (as valid species).

Atheta (s. str.) diffisa: Moore & Legner, 1975: 359   (as valid species).

Atheta (s. str.) nata: Moore & Legner, 1975: 367   (as valid species).

Atheta (s. str.) modiella: Moore & Legner, 1975: 366   (as valid species).

Atheta (s. str.) vacillans: Moore & Legner, 1975: 376   (as valid species).

Xenota frosti: Seevers, 1978: 268   (as valid species).

Xenota comitata: Seevers, 1978: 268   (as valid species).

Xenota gnoma: Seevers, 1978: 268   (as valid species).

Xenota elota: Seevers, 1978: 268   (as valid species).

Xenota insidiosa: Seevers, 1978: 268   (as valid species).

Pseudota puricula: Seevers, 1978: 260   (as valid species).

Xenota candidula: Seevers, 1978: 268   (as valid species).

Xenota diffisa: Seevers, 1978: 268   (as valid species).

Xenota nata: Seevers, 1978: 268   (as valid species).

Xenota modiella: Seevers, 1978: 268   (as valid species).

Xenota vacillans: Seevers, 1978: 269   (as valid species).

Type material. Lectotype of Atheta frosti   (here designated): ♀, “N.[ew] H.[ampshire] Fall”, “ frosti Brh. Cotypus   ”, “7597”, “Chicago NHMus M.Bernhauer Collection ”   . Paralectotypes: ♂, “Frmghm. [Framingham] Mass.[achusetts] Frost”, “ frosti Brh.   Typus” (yellow label), “7008”, “Chicago NHMus M.Bernhauer Collection ” ( FMNH)   ; ♀, “Penn.[sylvania]”, “Aug.[ust] [18]96”, red circle, “ nigritula Amer.   b. Gvh.”, “ frosti Brh. Cotypus   ” (yellow label), “Chicago NHMus M.Bernhauer Collection ”   ; ♂ (with the head and the apex of the abdomen missing), “Brooklyn, N.[ew] Y.[ork]”, “July”, white circle, “ frosti Brh. Cotypus   ” (yellow label), “Chicago NHMus M.Bernhauer Collection ” ( FMNH)   .

Lectotype of Atheta comitata   (here designated): ♂, “R.[hode] I.[sland, Boston Neck]”, “ Atheta comitata Csy.   ”, “TYPE USNM 39217” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: 3♂♂, 5♀♀, “R.[hode] I.[sland, Boston Neck]”, “ comitata   PARATYPE USNM 39217” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta gnoma   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ Atheta gnoma Csy.   ”, “TYPE USNM 39234” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta elota   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts.]”, “ Atheta elota Csy.   ”, “TYPE USNM 39219” (red label), “CASEY bequest 1925” ( NMNH). Paralectotypes: 4♂♂, 2♀♀, “N.[ew] Y.[ork, Catskill Mts.]”, “ elota   PARATYPE USNM 39219” (red label), “CASEY bequest 1925” ( NMNH) (one additional paralectotype of Atheta elota   (“N.[ew] Y.[ork, Catskill Mts.]”, “ elota   – 5 PARATYPE USNM 39219” (red label), “CASEY bequest 1925” ( NMNH)) belongs to Atheta klagesi Bernhauer   ).

Lectotype of Atheta insidiosa   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ Atheta insidiosa Csy.   ”, “TYPE USNM 39220” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: 7♂♂, 3♀♀, 1 specimen with the tip of the abdomen missing, “N.[ew] Y.[ork, Catskill Mts.]”, “ insidiosa   PARATYPE USNM 39220” (red label), “CASEY bequest 1925” ( NMNH) (9 additional paralectotypes (“N.[ew] Y.[ork, Catskill Mts. ]”, “ insidiosa   PARATYPE USNM 39220” (red label), “CASEY bequest 1925” ( NMNH) belong to Atheta klagesi Bernhauer   ).  

Lectotype of Pseudota puricula   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ puricula Csy.   ”, “TYPE USNM 39159” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotype: ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ puricula   PARATYPE USNM 39159” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta candidula   (here designated): ♀, “N.[ew] Y.[ork, Catskill Mts. ]”, “ candidula Csy.   ”, “TYPE USNM 39232” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta diffisa   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ diffisa Csy.   ”, “TYPE USNM 39233” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta nata   (here designated): ♀, “R.[hode] I.[sland, Boston Neck]”, “ nata Csy.   ”, “TYPE USNM 39228” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta modiella   (here designated): ♂, “R.[hode] I.[sland, Boston Neck]”, “ modicella [sic!] Csy.”, “TYPE USNM 39235” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Atheta vacillans   (here designated): ♀, “R.[hode] I.[sland, Boston Neck]”, “ vacillans Csy.   ”, “TYPE USNM 39222” (red label), “CASEY bequest 1925” ( NMNH).  

Additional material. UNITED STATES: North Carolina: Haywood Co.: ♂, 3♀♀, Cold Springs Creek, 29 km N Waynesville, 35º44.56'N 83º00.39'W, 650 m, 20.viii.1998 (V.I.Gusarov); Macon Co.: 3♂♂, ♀, Nantahala River, 26 km NWW Franklin, 35º15.37'N 83º39.24'W, 650 m, 21.viii.1998 (V.I.Gusarov) ( SPSU); Pennsylvania: ♂, near Philadelphia, ( NMNH (Casey collection)); Adams Co.: ♂, 3♀♀, 15 km E Chambersburg, Michaux State Forest, 39º54.87'N 77º28.33'W, 400 m, in mushrooms, 18.viii.1998 (V.I.Gusarov) ( SPSU); Vermont: Windsor Co.: ♀, 8 km SEE Springfield, Hwy. I­91, 43º14.00'N 72º26.81'W, 250 m, in forest litter, Acer   , Betula   , Tsuga   , Quercus   , 21.ix.1998 (V.I.Gusarov) ( SPSU).

Diagnosis. Atheta frosti   is closely related to Palaearctic At. nigritula ( Gravenhorst, 1802)   but differs in the shape of the male tergum 8 ( Figs. 70, 72 View FIGURES 70–74 ), the aedeagus ( Figs. 75– 79 View FIGURES 75–80 ) and spermatheca ( Fig. 80 View FIGURES 75–80 ).

Discussion. The types of At. frosti   , At. comitata   , At. gnoma   , At. elota   , At. insidiosa   , Ps. puricula   , At. candidula   , At. diffisa   , At. nata   , At. modiella   and At. vacillans   are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.

Distribution. Atheta frosti   is widely distributed in the eastern United States (Vermont, New Hampshire, Massachusetts, Rhode Island, New York, Pennsylvania and North Carolina) but its distribution is poorly documented.

FMNH

Field Museum of Natural History

NMNH

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Atheta

Loc

Atheta (Tetropla) frosti Bernhauer, 1909

Gusarov, Vladimir I. 2003
2003
Loc

Xenota frosti:

Seevers, C. H. 1978: 268
1978
Loc

Xenota comitata:

Seevers, C. H. 1978: 268
1978
Loc

Xenota gnoma:

Seevers, C. H. 1978: 268
1978
Loc

Xenota elota:

Seevers, C. H. 1978: 268
1978
Loc

Xenota insidiosa:

Seevers, C. H. 1978: 268
1978
Loc

Pseudota puricula: Seevers, 1978: 260

Seevers, C. H. 1978: 260
1978
Loc

Xenota candidula:

Seevers, C. H. 1978: 268
1978
Loc

Xenota diffisa:

Seevers, C. H. 1978: 268
1978
Loc

Xenota nata:

Seevers, C. H. 1978: 268
1978
Loc

Xenota modiella:

Seevers, C. H. 1978: 268
1978
Loc

Xenota vacillans:

Seevers, C. H. 1978: 269
1978
Loc

Atheta (s. str.) frosti:

Moore, I. & Legner, E. F. 1975: 361
1975
Loc

Atheta (s. str.) comitata: Moore & Legner, 1975: 357

Moore, I. & Legner, E. F. 1975: 357
1975
Loc

Atheta (s. str.) gnoma: Moore & Legner, 1975: 362

Moore, I. & Legner, E. F. 1975: 362
1975
Loc

Atheta (s. str.) elota: Moore & Legner, 1975: 360

Moore, I. & Legner, E. F. 1975: 360
1975
Loc

Atheta (s. str.) insidiosa: Moore & Legner, 1975: 363

Moore, I. & Legner, E. F. 1975: 363
1975
Loc

Atheta (Pancota) puricula:

Moore, I. & Legner, E. F. 1975: 371
Moore, I. & Legner, E. F. 1975: 356
1975
Loc

Atheta (s. str.) diffisa: Moore & Legner, 1975: 359

Moore, I. & Legner, E. F. 1975: 359
1975
Loc

Atheta (s. str.) nata: Moore & Legner, 1975: 367

Moore, I. & Legner, E. F. 1975: 367
1975
Loc

Atheta (s. str.) modiella: Moore & Legner, 1975: 366

Moore, I. & Legner, E. F. 1975: 366
1975
Loc

Atheta (s. str.) vacillans: Moore & Legner, 1975: 376

Moore, I. & Legner, E. F. 1975: 376
1975
Loc

Atheta (s. str.) frosti:

Bernhauer, M. & Scheerpeltz, O. 1926: 643
1926
Loc

Atheta (s. str.) comitata:

Bernhauer, M. & Scheerpeltz, O. 1926: 640
1926
Loc

Atheta (s. str.) gnoma:

Bernhauer, M. & Scheerpeltz, O. 1926: 643
1926
Loc

Atheta (s. str.) elota:

Bernhauer, M. & Scheerpeltz, O. 1926: 642
1926
Loc

Atheta (s. str.) insidiosa:

Bernhauer, M. & Scheerpeltz, O. 1926: 644
1926
Loc

Atheta (Pancota) puricula:

Bernhauer, M. & Scheerpeltz, O. 1926: 661
1926
Loc

Atheta (s. str.) candidula:

Bernhauer, M. & Scheerpeltz, O. 1926: 639
1926
Loc

Atheta (s. str.) diffisa:

Bernhauer, M. & Scheerpeltz, O. 1926: 641
1926
Loc

Atheta (s. str.) nata:

Bernhauer, M. & Scheerpeltz, O. 1926: 646
1926
Loc

Atheta (s. str.) modiella:

Bernhauer, M. & Scheerpeltz, O. 1926: 645
1926
Loc

Atheta (s. str.) vacillans:

Bernhauer, M. & Scheerpeltz, O. 1926: 651
1926
Loc

Atheta (s. str.) frosti:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) comitata:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) gnoma:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) elota:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) insidiosa:

Fenyes, A. 1920: 213
1920
Loc

Atheta (Pancota) puricula:

Fenyes, A. 1920: 202
1920
Loc

Atheta (s. str.) candidula:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) diffisa:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) nata:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) modiella:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) vacillans:

Fenyes, A. 1920: 214
1920
Loc

Pseudota puricula

Casey, T. L. 1911: 148
1911
Loc

Atheta (s. str.) candidula

Casey, T. L. 1911: 94
1911
Loc

Atheta (s. str.) diffisa

Casey, T. L. 1911: 95
1911
Loc

Atheta (s. str.) nata

Casey, T. L. 1911: 95
1911
Loc

Atheta (s. str.) modiella

Casey, T. L. 1911: 96
1911
Loc

Atheta (s. str.) vacillans

Casey, T. L. 1911: 98
1911
Loc

Atheta (s. str.) comitata

Casey, T. L. 1910: 24
1910
Loc

Atheta (s. str.) gnoma

Casey, T. L. 1910: 25
1910
Loc

Atheta (s. str.) elota

Casey, T. L. 1910: 26
1910
Loc

Atheta (s. str.) insidiosa

Casey, T. L. 1910: 26
1910
Loc

Atheta (s. str.) frosti

Bernhauer, M. 1909: 520
1909
Loc

Atheta (s. str.) nigritula:

Bernhauer, M. 1907: 392
1907