Atheta klagesi Bernhauer, 1909

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134: 66-70

publication ID

http://doi.org/ 10.11646/zootaxa.353.1.1

publication LSID

lsid:zoobank.org:pub:104E9C0B-B207-41E6-816C-E91FC1C7DBA0

DOI

http://doi.org/10.5281/zenodo.5102970

persistent identifier

http://treatment.plazi.org/id/03F387B2-FFC1-5309-CD24-5746FB0DA361

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scientific name

Atheta klagesi Bernhauer, 1909
status

 

Atheta klagesi Bernhauer, 1909  

( Figs. 93–102 View FIGURES 93–97 View FIGURES 98–102 )

Atheta (s. str.) klagesi Bernhauer, 1909: 524   .

Atheta (Delphota) cephalina Casey, 1910a: 18   , syn. nov.

Atheta (s. str.) nympha Casey, 1910a: 42   , syn. nov.

Atheta (s. str.) discreta Casey, 1910a: 42   (nec Casey, 1893: 335, nec Casey, 1910a: 79), syn. nov.

Pseudota dissensa Casey, 1910a: 115   , syn. nov.

Atheta (s. str.) villica Casey, 1911a: 97   , syn. nov.

Atheta (s. str.) klagesi: Fenyes, 1920: 213   (as valid species).

Atheta (s. str.) cephalina: Fenyes, 1920: 215   (as valid species).

Atheta (s. str.) nympha: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) discreta: Fenyes, 1920: 214   (as preoccupied name).

Atheta (Pancota) dissensa: Fenyes, 1920: 202   (as valid species).

Atheta (s. str.) villica: Fenyes, 1920: 214   (as valid species).

Atheta (s. str.) klagesi: Bernhauer & Scheerpeltz, 1926: 644   (as valid species).

Atheta (s. str.) cephalina: Bernhauer & Scheerpeltz, 1926: 640   (as valid species).

Atheta (s. str.) nympha: Bernhauer & Scheerpeltz, 1926: 647   (as valid species).

Atheta (s. str.) discreta: Bernhauer & Scheerpeltz, 1926: 641   (as preoccupied name).

Atheta (Pancota) dissensa: Bernhauer & Scheerpeltz, 1926: 661   (as valid species).

Atheta (s. str.) villica: Bernhauer & Scheerpeltz, 1926: 651   (as valid species).

Atheta (s. str.) disca Moore & Legner, 1975: 359   (replacement name for At. discreta Casey, 1910a: 42   ), syn. nov.

Atheta (s. str.) discreta: Moore & Legner, 1975: 359   (as synonym of At. disca   ).

Xenota klagesi: Seevers, 1978: 268   (as valid species).

Xenota cephalina: Seevers, 1978: 268   (as valid species).

Xenota nympha: Seevers, 1978: 268   (as valid species).

Xenota discreta: Seevers, 1978: 268   (as valid species).

Pseudota dissensa: Seevers, 1978: 260   (as valid species).

Xenota villica: Seevers, 1978: 269   (as valid species).

Type material. Lectotype of Atheta klagesi   (here designated): ♂, “Me. [Maine]”, “ Frost ”, “1654.”, “153.”, “ Klagesi Brh. Cotypus. Fenyes” (yellow label), “ Chicago NHMus M.Bernhauer Collection ” (FMNH). Paralectotypes: ♀, “Jeannette, viii, Pa. [Pennsylvania], “Klages”, “ Klagesi Brh. Cotypus. Fenyes” (yellow label), “Chicago NHMus M.Bernhauer Collection ”; ♂, “Jeannette, viii, Pa. [Pennsylvania], H.G.Klages. ”, “ Klagesi Brh. Typus. Fenyes” (yellow label), “Chicago NHMus M.Bernhauer Collection ” (FMNH).

Lectotype of Atheta cephalina   (here designated): ♂, “Ia. [Iowa ( Brendel )]”, “ ♂ ”, “ Delphota cephalina Csy.   ”, “TYPE USNM 39485” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotype: ♀, “Ia. [Iowa ( Brendel )]”, “ cephalina PARATYPE USNM 39485” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta nympha   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ Atheta nympha Csy.   ”, “TYPE USNM 39276” (red label), “CASEY bequest 1925” ( NMNH)   . Paralectotypes: 5♂♂, 3♀♀, “N.[ew] Y.[ork, Catskill Mts. ]”, “ nympha   PARATYPE USNM 39276” (red label), “CASEY bequest 1925” ( NMNH)   .

Lectotype of Atheta discreta   (here designated): ♂, “N.[ew] Y.[ork, Catskill Mts. ]”, “ Atheta discreta Csy.   ”, “TYPE USNM 39278” (red label), “CASEY bequest 1925” ( NMNH).  

Lectotype of Pseudota dissensa   (here designated): ♂, “Penn.[sylvania, Philadelphia]”, “ Pseudota dissensa Csy.   ”, “TYPE USNM 39147” (red label), “CASEY bequest 1925” ( NMNH).  

Holotype of Atheta villica   : ♀, “Penn.[sylvania, near Philadelphia]”, “ villica Csy.   ”, “TYPE USNM 39231” (red label), “CASEY bequest 1925” ( NMNH).  

Additional material. UNITED STATES: Minnesota: Lake Co.: ♂, 22 km NE Two Harbors, Split Rock Lighthouse State Park, Hwy. 61, 47º10.99'N 91º24.52'W, 70 m, in forest litter, Betula   , Picea   , Populus   , Rubus   , Majanthemum   , ferns, 10.vii.1999 (V.I.Gusarov) ( SPSU); New Jersey: Warren Co.: 2♀♀, 24 km NW Hackettstown, Dunnfield Creek Nat. Area, 40º58.33'N 75º07.49'W, 20 m, Quercus   , Acer   , 5.iv.1999 (V.I.Gusarov) ( SPSU); New York: Orange Co.: 3♂♂, 2♀♀, 8 km S New Windsor, Black Rock Forest, 9.v.1998 (V.I.Gusarov); 5♂♂, 2♀♀, ditto but 23–25.v.1998 ( SPSU); CANADA: Québec: ♂, Parc de la Gatineau, 11 km E Chelsea, Champlain, 26.ix.1998 (V.I.Gusarov); ♂, Chûte­aux­Galets, 1.viii.1994; ♂, 190 km NW Montréal, Mont­Laurier, 46º34'N 75º29'W, 5.vii.1994 ( SPSU).

Diagnosis. Atheta klagesi   can be recognized by the following combination of characters: head and abdominal terga 5–6 black; pronotum, abdominal terga 3–4 and 7–8, and antennal articles 4–11 or 3–11 brown; elytra yellow with vaguely defined darker areas around scutellum and in postero­lateral angles; antennal articles 1–2 or 1–3, legs and mouthparts yellow; antennal articles 5–10 strongly transverse, article 10 twice as wide as long; pronotal pubescence of type V ( Benick & Lohse 1974); mesotibial macrosetae thin, as long as tibial width; posterior margin of male tergum 8 convex and crenulate ( Figs. 93, 95 View FIGURES 93–97 ); aedeagus as in Figs. 99–102 View FIGURES 98–102 ; and spermatheca as in Fig. 98 View FIGURES 98–102 .

Discussion. The types of At. klagesi   , At. cephalina   , At. nympha   , At. discreta   , Ps. dissensa   and At. villica   are similar in external characters and the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.

Atheta cephalina   is the type species of the subgenus Delphota Casey, 1910a   by original designation. Pseudota dissensa   is the type species of the genus Pseudota Casey, 1910a   by original designation (Casey first species rule). Casey did not specifically state in his description of Pseudota   that Ps. dissensa   was the type species of Pseudota   ; however, while designating the type species of Noverota Casey, 1910a   in the same paper, Casey stated (1910a, p. 90): “The first species may be regarded as the type, as in all other cases where the type is not specifically named”.

Since both At. cephalina   and Ps. dissensa   are synonyms of At. klagesi   , the names Delphota   and Pseudota   are subjective synonyms of each other, irrespective of what subgenera in the genus Atheta   are accepted as valid. The problem of subgeneric placement of At. klagesi   within Atheta   is outside the scope of the present paper. The matter will be addressed in a separate work when the generic revision of the tribe Athetini   is completed.

Distribution. Atheta klagesi   is known from the eastern United States (Minnesota, Iowa, Pennsylvania, New Jersey, New York, Maine) and eastern Canada (Québec).

NMNH

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Atheta

Loc

Atheta klagesi Bernhauer, 1909

Gusarov, Vladimir I. 2003
2003
Loc

Xenota klagesi:

Seevers, C. H. 1978: 268
1978
Loc

Xenota cephalina:

Seevers, C. H. 1978: 268
1978
Loc

Xenota nympha:

Seevers, C. H. 1978: 268
1978
Loc

Xenota discreta:

Seevers, C. H. 1978: 268
1978
Loc

Pseudota dissensa: Seevers, 1978: 260

Seevers, C. H. 1978: 260
1978
Loc

Xenota villica:

Seevers, C. H. 1978: 269
1978
Loc

Atheta (s. str.) disca

Moore, I. & Legner, E. F. 1975: 359
Casey, T. L. 1910: 42
1975
Loc

Atheta (s. str.) discreta: Moore & Legner, 1975: 359

Moore, I. & Legner, E. F. 1975: 359
1975
Loc

Atheta (s. str.) klagesi: Bernhauer & Scheerpeltz, 1926: 644

Bernhauer, M. & Scheerpeltz, O. 1926: 644
1926
Loc

Atheta (s. str.) cephalina: Bernhauer & Scheerpeltz, 1926: 640

Bernhauer, M. & Scheerpeltz, O. 1926: 640
1926
Loc

Atheta (s. str.) nympha: Bernhauer & Scheerpeltz, 1926: 647

Bernhauer, M. & Scheerpeltz, O. 1926: 647
1926
Loc

Atheta (s. str.) discreta:

Bernhauer, M. & Scheerpeltz, O. 1926: 641
1926
Loc

Atheta (Pancota) dissensa:

Bernhauer, M. & Scheerpeltz, O. 1926: 661
1926
Loc

Atheta (s. str.) villica: Bernhauer & Scheerpeltz, 1926: 651

Bernhauer, M. & Scheerpeltz, O. 1926: 651
1926
Loc

Atheta (s. str.) klagesi:

Fenyes, A. 1920: 213
1920
Loc

Atheta (s. str.) cephalina:

Fenyes, A. 1920: 215
1920
Loc

Atheta (s. str.) nympha:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) discreta:

Fenyes, A. 1920: 214
1920
Loc

Atheta (Pancota) dissensa:

Fenyes, A. 1920: 202
1920
Loc

Atheta (s. str.) villica:

Fenyes, A. 1920: 214
1920
Loc

Atheta (s. str.) villica

Casey, T. L. 1911: 97
1911
Loc

Atheta (Delphota) cephalina

Casey, T. L. 1910: 18
1910
Loc

Atheta (s. str.) nympha

Casey, T. L. 1910: 42
1910
Loc

Atheta (s. str.) discreta

Casey, T. L. 1910: 42
Casey, T. L. 1910: 79
Casey, T. L. 1893: 335
1910
Loc

Pseudota dissensa

Casey, T. L. 1910: 115
1910
Loc

Atheta (s. str.) klagesi

Bernhauer, M. 1909: 524
1909