Strigota ambigua ( Erichson, 1839 ) Gusarov, 2003

Gusarov, Vladimir I., 2003, Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes, Zootaxa 353 (1), pp. 1-134 : 107-109

Strigota ambigua ( Erichson, 1839) , comb. nov.

( Figs. 142–154 )

Homalota ambigua Erichson, 1839: 134 .

Strigota oppidana Casey, 1910a: 177 , syn. nov.

Strigota gnava Casey, 1910a: 177 , syn. nov.

Strigota verecunda Casey, 1910a: 178 , syn. nov.

Strigota assueta Casey, 1910a: 178 , syn. nov.

Strigota mediocris Casey, 1910a: 179 , syn. nov.

Strigota vapida Casey, 1910a: 179 , syn. nov.

Strigota inculta Casey, 1910a: 180 , syn. nov.

Strigota placata Casey, 1910a: 182 , syn. nov.

Strigota (s. str.) recta Casey, 1911a: 165 , syn. nov.

Paradilacra ambigua: Fenyes, 1920: 244 (as valid species).

Strigota oppidana: Fenyes, 1920: 253 (as valid species).

Strigota gnava: Fenyes, 1920: 253 (as valid species).

Strigota verecunda: Fenyes, 1920: 253 (as valid species).

Strigota assueta: Fenyes, 1920: 253 (as valid species).

Strigota mediocris: Fenyes, 1920: 253 (as valid species).

Strigota vapida: Fenyes, 1920: 253 (as valid species).

Strigota inculta: Fenyes, 1920: 253 (as valid species).

Strigota placata: Fenyes, 1920: 253 (as valid species).

Strigota recta: Fenyes, 1920: 253 (as valid species).

Anaduosternum brevipennis Notman, 1922: 107 , syn. nov.

Atheta (Paradilacra) ambigua: Bernhauer & Scheerpeltz, 1926: 607 (as valid species).

Strigota oppidana: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota gnava: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota verecunda: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota assueta: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota mediocris: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota vapida: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota inculta: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota placata: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Strigota recta: Bernhauer & Scheerpeltz, 1926: 590 (as valid species).

Atheta (Anaduosternum) brevipennis: Bernhauer & Scheerpeltz, 1926: 671 (as valid species).

Atheta (Dilacra) ambigua: Moore & Legner, 1975: 354 (as valid species).

Strigota oppidana: Moore & Legner, 1975: 486 (as valid species).

Strigota gnava: Moore & Legner, 1975: 486 (as valid species).

Strigota verecunda: Moore & Legner, 1975: 486 (as valid species).

Strigota assueta: Moore & Legner, 1975: 486 (as valid species).

Strigota mediocris: Moore & Legner, 1975: 486 (as valid species).

Strigota vapida: Moore & Legner, 1975: 486 (as valid species).

Strigota inculta: Moore & Legner, 1975: 486 (as valid species).

Strigota placata: Moore & Legner, 1975: 486 (as valid species).

Strigota recta: Moore & Legner, 1975: 486 (as valid species).

Atheta (Anaduosternum) notmani Moore & Legner, 1975: 367 (replacement name for At. brevipennis ( Notman, 1922) (nec J. Sahlberg, 1876)), syn. nov.

Atheta (Anaduosternum) brevipennis: Moore & Legner, 1975: 367 (as synonym of At. notmani ).

Paradilacra ambigua: Seevers, 1978: 265 (as valid species).

Strigota oppidana: Seevers, 1978: 258 (as valid species).

Strigota gnava: Seevers, 1978: 257 (as valid species).

Strigota verecunda: Seevers, 1978: 258 (as valid species).

Strigota assueta: Seevers, 1978: 257 (as valid species).

Strigota mediocris: Seevers, 1978: 257 (as valid species).

Strigota vapida: Seevers, 1978: 258 (as valid species).

Strigota inculta: Seevers, 1978: 258 (as valid species).

Strigota placata: Seevers, 1978: 258 (as valid species).

Strigota recta: Seevers, 1978: 258 (as valid species).

Anaduosternum brevipennis: Seevers, 1978: 261 (as valid species).

Type material. Lectotype of Homalota ambigua (here designated): ♀, “5507”, “ ambigua Er. Am. [erica] spt. Zimm.[ermann]” (green label), “Zool. Mus. Berlin ”, “Typus” (red label), “ Homalota ambigua Er. [on red side] / Lectotypus Lohse fix. 1983 [on white side]” [ Lohse never published this lectotype designation] ( ZMHB) . Paralectotypes: 2♂♂, 2♀♀, “America Sept. Zimmermann Nr. 5507” (green label), “Zool. Mus. Berlin ”, “Typus” (red label) ( ZMHB) .

Lectotype of Strigota oppidana (here designated): ♀, “N.[ew] Y.[ork, near New York City ]”, “ Strigota oppidana Csy. ”, “TYPE USNM 39034” (red label), “CASEY bequest 1925” ( NMNH) . Paralectotypes: 2♂♂, 2♀♀, “N.[ew] Y.[ork, near New York City ]”, “ oppidana PARATYPE USNM 39034” (red label), “CASEY bequest 1925” ( NMNH) .

Lectotype of Strigota gnava (here designated): ♂, “·N.[orth] C.[arolina, Asheville]”, “ gnava Csy. ”, “TYPE USNM 39035” (red label), “CASEY bequest 1925” ( NMNH). Paralectotypes: ♂, ♀, “·N.[orth] C.[arolina, Asheville]”, “ gnava PARATYPE USNM 39035” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Strigota verecunda (here designated): ♂, “Ia. [Iowa, Cedar Rapids ( Brendel )]”, “ verecunda Csy. ”, “TYPE USNM 39037” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Strigota assueta (here designated): ♀, “Mo.· [Missouri, St. Louis ]”, “ assueta Csy. ”, “TYPE USNM 39038” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Strigota mediocris (here designated): ♀, “·Tex.[as, El Paso]”, “ mediocris Csy. ”, “TYPE USNM 39039” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Strigota vapida (here designated): ♂, “Salida Colo[rado] Wickham”, “ vapida ­3 PARATYPE USNM 39040” (red label), “CASEY bequest 1925” ( NMNH). Paralectotype: ♂, “Cöl. [Colorado, Greeley]”, “ vapida ­2 PARATYPE USNM 39040” (red label), “CASEY bequest 1925” (the second paralectotype of St. vapida (♂, “Coolidge N.[ew] Mex.[ico]”, “ vapida Csy. ”, “TYPE USNM 39040” (red label), “CASEY bequest 1925” ( NMNH)) is not conspecific with St. vapida and belongs to St. seducens Casey, 1910a or St. perplexa Casey, 1910a ).

Lectotype of Strigota inculta (here designated): ♀, “·Nev.[ada, Reno]”, “ inculta Csy. ”, “TYPE USNM 39041” (red label), “CASEY bequest 1925” ( NMNH). Paralectotypes: 2♂♂, ♀, “·Nev.[ada, Reno]”, “ inculta PARATYPE USNM 39041” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Strigota placata (here designated): ♂, “Cal.[ifornia, Santa Cruz]”, “ placata Csy. ”, “TYPE USNM 39045” (red label), “CASEY bequest 1925” ( NMNH). Paralectotype: ♂, “Cal.[ifornia, Santa Cruz]”, “ placata PARATYPE USNM 39045” (red label), “CASEY bequest 1925” ( NMNH).

Lectotype of Strigota recta (here designated): ♀, “ West Haven Ct [Connecticut] 27 June 1905 H.L.Viereck ”, “ recta Csy. ”, “TYPE USNM 39036” (red label), “CASEY bequest 1925” ( NMNH).

Holotype of Anaduosternum brevipennis : UNITED STATES: New Jersey: ♂, Newark, 24.vii (E.L.Dickerson) ( SIIS) . Paratype: UNITED STATES: New Jersey: ♂, Newark, 24.vii (E.L.Dickerson) ( SIIS) .

Additional material. UNITED STATES: California: Kern Co.: ♂, ♀, 4 specimens, Lake Isabella, Rd. 155, 35º38.37'N 118º29'W, 800 m, Isabella Lake Main Dam, river bank below dam, 12.viii.1999 (V.I.Gusarov) ( KSEM) ; Kansas: Douglas Co.: 10 specimens, 2.5 km N Lawrence, flood refuse by the Kansas River (right bank), 38º59.59'N 95º14.59'W, 200 m, 18.iv.1999 (V.I.Gusarov) ( SPSU) ; New York: 2♀♀, “L.[ong] I.[sland]” ( NMNH (Casey Collection )) ; Texas: Brewster Co.: 2♀♀, 19 km S Study Butte, 29º09.41'N 103º33.39'W, 650 m, Rio Grande , river banks, 31.iii.2001 (V.I.Gusarov) ( SPSU) .

Diagnosis. Strigota ambigua is very similar to other Nearctic species of Strigota and can be reliably identified only by the distinct shape of the aedeagus ( Figs. 149–154 ) and spermatheca ( Figs. 147–148 ).

Discussion. The types of H. ambigua , St. oppidana , St. gnava , St. verecunda , St. assueta , St. mediocris , St. vapida , St. inculta , St. placata , St. recta and Ana. brevipennis are similar in external characters and in the shape of the aedeagus and spermatheca. All these names are considered to be synonyms.

Moore and Legner (1975) proposed the name Atheta notmani to replace the name Atheta brevipennis ( Notman, 1922) (ex Anaduosternum ) which was preoccupied by the name Atheta brevipennis (J. Sahlberg, 1876) (ex Homalota ), listed by Bernhauer & Scheerpeltz (1926) as a junior synonym of Atheta (Anopleta) arcana ( Erichson, 1839) . Atheta notmani is a junior synonym of Strigota ambigua .

Distribution. Strigota ambigua is widespread across the United States. It is known from New York, Connecticut, New Jersey, North Carolina, Iowa, Missouri, Kansas, Texas, Colorado, New Mexico, Nevada and California.

Figures

FIGURES 142–146. Abdominal segment 8 of Strigota ambigua (Erichson) (male, paralectotype of St. gnava Casey (142–144); and female, lectotype of St. oppidana Casey (145–146)). 142 — male tergum 8; 143 — male sternum 8; 144 — apex of male sternum 8, dorsal [sic!] view; 145 — female tergum 8; 146 — female sternum 8. Scale bar 0.1 mm (144), 0.2 mm (142–143, 145–146).

FIGURES 147–154. Genitalia of Strigota ambigua (Erichson) (male, paralectotype of St. gnava Casey (149–154); and female, lectotype of St. oppidana Casey (147–148)). 147–148 — spermatheca (of the same specimen; different views); 149 — median lobe, parameral view; 150 — apex of median lobe, parameral view; 151 — median lobe, lateral view; 152 — apex of median lobe, lateral view; 153–154 — apex of median lobe with retracted internal sac, lateral view. Scale bar 0.1 mm (150, 152), 0.2 mm (147–149, 151, 153–154).