Deltochilum (Deltohyboma) spinipes Paulian, 1938

Silva, Fernando A. B. & Vaz-De-Mello, Fernando Z., 2014, A new giant species of Deltochilum subgenus Deltohyboma (Coleoptera, Scarabaeidae, Scarabaeinae) from Colombia, with notes on D. spinipes Paulian, 1938, Zootaxa 3802 (2), pp. 276-284 : 281-283

Deltochilum (Deltohyboma) spinipes Paulian, 1938

( Fig. 3 )

Paulian, 1938: 280

Notes. Paulian described this species based on several specimens from his collection as well as the collections of Oberthür, Boucomont, and the British Museum. He designated a pair of specimens from the Oberthür collection as " types " and the others as "cotypes"", but there was no formal Holotype designation and all type material is therefore syntypic. All syntypes in MNHN of D. spinipes have a body length less than 20 mm. However, Paulian states the length for the species as " 18-24 mm ". Based on this fact, we believe that the 24 mm female found in a separate box (female of D. gigante , see above) was regarded by Paulian as spinipes while describing this species.

The type series of spinipes shows variation and we strongly believe it to include more than one species. Therefore, in order to maintain stability of nomenclature, a lectotype is herein designated.

Two of the paralectotypes (♀4 and ♀5) appear to be non-conspecific with the remainder of the type series and likely represent two different species. They differ in being blue (not bright green), much darker and dull; one specimen is bigger and duller than the other; both specimens were collected in Ecuador (not Colombia as the remainder of the type series). These two specimens, at a first glance, most likely belong to the species-complex that includes D. aequinoctiale and D. speciosissimum . However, we cannot be sure about the identity of D. aequinoctiale since its type has not yet been located (see Génier 2001).

Material examined. Lectotype (here designated): ♂. (old white label, handwritten). Bogota. / 2 (old white label, printed and bordered in black). Ex-Musaeo Mniszech./ 3 (white label, handwritten). D. spinipes n. sp. Type./ 4 (white label, bordered in black) R. PAULIAN Vidit./ 5 (white label, printed in black). MUSÉUM PARIS 1952 coll. R. OBERTHÜR./ 6 (red label, printed in black) SYNTYPE./ 7 (white label, printed in black) MNHN EC 2509./ 8 (red label, printed in black) LECTOTYPE./ 9 (white label, handwritten, printed and bordered in black) Deltochilum spinipes Paulian LECTO. F Vaz-de-Mello des. 2013. Figure 3 A (copyright MNHN /A. Mantilleri). Paralectotypes: 2 ♂♂, 5 ♀♀ ( MNHN):

♂1. 1 (old white label, printed and bordered in black). COLOMBIE COLLECTION LE MOULT. / 2 (white label, handwritten and printed in black). D. spinipes Paul. R. Paulian det./ 3 (white label, printed in black). MUSÉUM PARIS 1952 coll. R. OBERTHÜR./ 4 (red label, printed in black) SYNTYPE./ 5 (white label, printed in black) MNHN EC 2508./ 6 (yellow label, printed in black) PARALECTOTYPE./ 7 (white label, handwritten, printed and bordered in black) Deltochilum spinipes Paulian PARALECTO. F Vaz-de-Mello des. 2013. Figures 3 B, C, F and G (copyright MNHN /A. Mantilleri).

♂2. 1 (old white label, handwritten). Fusagasugá./ 2 (old white label, handwritten). pretiosum Harold./ 3 (old white label, printed and bordered in black). Ex-Musaeo E. Steinheil./ 4 (white label, bordered in black) R. PAULIAN Vidit./ 5 (yellow label, printed in black) PARALECTOTYPE./ 6 (white label, handwritten, printed and bordered in black) Deltochilum spinipes Paulian PARALECTO. F Vaz-de-Mello des. 2013.

♀1. 1 (old white label, printed and bordered in black). COLOMBIE ANTIOQUIA./ 2 (old white label, printed and bordered in black) COLLON LE MOULT Naturaliste Paris. / 3 (white label, bordered in black) R. PAULIAN Vidit./ 4 (white label, printed in black). MUSÉUM PARIS 1952 coll. R. OBERTHÜR./ 5 (white label, printed in black) Deltochilum spinipes Paulian. / 6 (red label, printed in black) SYNTYPE./ 7 (white label, printed in black) MNHN EC 2510./ 8 (yellow label, printed in black) PARALECTOTYPE./ 9 (white label, handwritten, and printed and bordered in black) Deltochilum spinipes Paulian PARALECTO. F Vaz-de-Mello des. 2013. Figures 3 D and E (copyright MNHN /A. Mantilleri).

♀2. 1 (old white label, handwritten). Muzo, Columbia./ 2 (old white label, printed and bordered in black). Ex- Musaeo D. Sharp 1890./ 3 (white label, bordered in black) R. PAULIAN Vidit./ 4 (yellow label, printed in black) PARALECTOTYPE./ 5 (white label, handwritten, printed and bordered in black) Deltochilum spinipes Paulian PARALECTO. F Vaz-de-Mello des. 2013.

♀3. 1 (old white label, printed and bordered in black). COLOMBIE COLLECTION LE MOULT. / 2 (white label, bordered in black) R. PAULIAN Vidit./ 3 (yellow label, printed in black) PARALECTOTYPE./ 4 (white label, handwritten, printed and bordered in black) Deltochilum spinipes Paulian PARALECTO. F Vaz-de-Mello des. 2013.

♀4. 1 (white label, printed and bordered in black) Equateur Env. d'Ambato R.P. Irenée Blanc./ 2 (white label, bordered in black) R. PAULIAN Vidit./ 3 (yellow label, printed in black) PARALECTOTYPE./ 4 (white label, handwritten, printed and bordered in black) Deltochilum spinipes Paulian PARALECTO. F Vaz-de-Mello des. 2013./ 5 (white label, handwritten, printed and bordered in black) DIFFERENT of D spinipes lectot. F Vaz-de- Mello des. 2013.

♀5. 1 (old white label, handwritten) Ecuador Buckley./ 2 (white label, printed and bordered in black) Ex- Musaeo H.W.Bates 1892./ 3 (white label, bordered in black) R. PAULIAN Vidit./ 4 (yellow label, printed in black) PARALECTOTYPE./ 5 (white label, handwritten, printed and bordered in black) Deltochilum spinipes Paulian PARALECTO. F Vaz-de-Mello des. 2013./ 6 (white label, handwritten, printed and bordered in black) DIFFERENT of D spinipes lectot. F Vaz-de-Mello des. 2013.

Figures

FIGURE 3. Deltochilum spinipes. (A) lectotype ♂, dorsal view; (B) paralectotype ♂ 1, dorsal view; (C) paralectotype ♂ 1, ventral view; (D) paralectotype ♀ 1, dorsal view; (E) paralectotype ♀ 1, ventral view; (F) paralectotype ♂ 1, aedeagus (lateral view); (G) paralectotype ♂ 1, aedeagus (dorsal view). Photographic copyright A. Mantilleri (MNHN).