Spogostylum griseipenne ( Macquart, 1850 )

Greathead, D. J., 2003, A sympatric species pair: Spogostylum ocyale (Wiedemann, 1828) and S. griseipenne Macquart, 1850 (Diptera: Bombyliidae), Zootaxa 274, pp. 1-6 : 5

Spogostylum griseipenne ( Macquart, 1850)

Exoprosopa griseipennis Macquart, 1850: 408 ( Egypt)

Argyramoeba ocyalis (Wiedemann) , François, 1972: 21 ( Mauritania) Spogostylum sordidum Sack, Greathead, 1980:230 ( Saudi Arabia males) Spogostylum ocyale (Wiedemann) , Zaitzev, 1997: 911 ( Israel)

Theodor (1983: Figs 598­599) drew attention to the existence of a second species near S. ocyale with different genitalia. Zaitzev (1977) disagreed and designated this species as S. ocyale . It had first been illustrated by François (1972: Figs 41­42) who noted that the male genitalia were different to those illustrated for S. ocyale by Greathead (1967: Fig. 50) and also provided a figure of the wing ( Fig. 4 ). Greathead (1980b: Fig. 19) illustrated the male genitalia as those of S. sordidum .

Diagnosis. Sides of interradial vein with a dark infuscation. Male with aedeagus longer and widened at the apex ( Theodor, 1983: François, 1972: Fig. 41; Fig. 598: Greathead, 1980b: Fig. 19, as S. sordidum ). Female with spermathecal bulb almost spherical ( Theodor, 1983: Fig. 597). Specimens tend towards the dark end of the range of coloration.

Distribution. Afrotropical: Mauritania ( François, 1972), Niger. Palaearctic: Egypt, Iraq, Libya, ‘Palestine’, Saudi Arabia, Tunisia, United Arab Emirates.

Material examined. Afrotropical. NIGER: In Zinkad, Tamesna, 25.vi.1990, A. Foucart, 1 male in DJG.

Palaearctic. EGYPT: 2 male syntypes, in UMO; Siwa, 12.vii.1935, J. Omer­Cooper, 1 male, BMNH; Kharga, 7.xi.1997, Magdi, 1 male in DJG. IRAQ: Tigris, Daurah, vvi.1920, A.D. Fraser, 1 male, 2 females in BMNH; Daurah, v.1920, A.D. Fraser, 1 male in BMNH; R. Dryala, 9.vii.1918, H.B. Peile, 1 male, 1 female in BMNH. JORDAN: Irbid Gov., Al Adasiyah, 32 40N 35 37E, 21.ix.1995, 1 male; Ma'an Gov., Ad Dinah, 29 37N 35 33E, 20.vii.1996, 1 male. LIBYA: N shore L. Karun, 20.ix.1945, R.L. Coe, 2 males in BMNH; Cyrenaica, Gambut, Wadi El Mra, 9.viii.1957, K.M. Guichard, 1 male in BMNH. ‘ PALESTINE ’: Galilee, 1924, O. Theodor, I male, 1 female in BMNH; Jericho, ixx.1918, O. Theodor, 1 male in BMNH. SAUDI ARABIA: Taif escarpment, 29.ix.1982, A.R. Pittaway, 1 male in BMNH; same, 6.xi.1982, 1 female; Khulays, 15.iv.1971, Locust Research Station, 1 male in DJG; Jedda, Bahara, 15.iv.1980, K.M. Guichard, 2 males in DJG. TUNISIA: Tozeur, 2.viii.1978, K.M. Guichard, 1 male in DJG. UNITED ARAB EMIRATES: Sharjah, 1969, D. Burgess, 1 male in BMNH; same, 4.xii.1970, M.D. Gallaher, 1 male in BMNH.

Figures

FIGURES 3 ­ 6. Genitalia of Spogostylum species. 3. S. ocyale (Wiedemann), phallosome. 4. S. griseipenne (Macquart), phallosome. 5. S. ocyale, bulb of spermatheca. 6. S. griseipenne, bulb of spermatheca. Scale bars = 0.2 mm.