Humptulipsia nobuharai

Kaim, Andrzej, Jenkins, Robert G., Tanabe, Kazushige & Kiel, Steffen, 2014, Mollusks from late Mesozoic seep deposits, chiefly in California, Zootaxa 3861 (5), pp. 401-440: 417-418

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Humptulipsia nobuharai

sp. nov.

Humptulipsia nobuharai   sp. nov.

( Fig. 7 View FIGURE 7. A – D G, J –M)

Type locality and strata: Sada Limestone, Shikoku, Japan; Campanian-Maastrichian (Late Cretaceous); for details see Nobuhara et al. (2008).

Holotype: UMUT MM 31170, two moderately well-preserved whorls 26.8 mm high and 15.4 mm wide.

Paratypes: UMUT MM 31230, adult specimen, UMUT MM 31171, UMUT MM 31172, two juvenile shells, UMUT MM 31173 fragmentary specimen displaying the humptulipsid slit on the shell base.

Additional material: Many unprepared specimens from the type locality. This gastropod occurs in mass accumulations at the type locality.

Diagnosis. Tall shell with largest whorl diameter in mid-flank. Adult shell smooth apart from prosocyrt growth. Shell base with narrow slit in its abapical portion.

Description. Tall shell with largest whorl diameter at mid-whorl. Protoconch and juvenile teleoconch unknown. Adult shell smooth apart from prosocyrt growth lines. Shell base with narrow slit in its abapical portion. Suture weakly incised. Aperture poorly preserved, drop-shaped.

Remarks. Humptulipsia nobuharai   is the third species of Humptulipsia described so far. The type species H. raui   is known from Eocene/Oligocene hydrocarbon seeps in Washington state (Kiel 2008) and differs from H. nobuharai   in having a much wider slit that is located much higher on the whorls. Humptulipsia macsotayi Kiel et al., 2010   described from Hauterivian (Early Cretaceous) presumed seep carbonates in southeastern France (Kiel et al. 2010) differs from H. nobuharai   by being ornamented throughout the entire shell ontogeny. We include Humptulipsia in the Hokkaidoconchidae due to the hokkaidoconchid-like ornamentation of the juvenile shell in H. raui   (see Fig. 1.9 in Kiel 2008 and Fig. 7 View FIGURE 7. A – D H herein). A slit similar to that of Humptulipsia is present in the members of family Pseudolividae   de Gregorio, 1880, which differ, however, in ornamentation style of the early whorls (see e.g., Pacaud & Schnetler 1999) and are known in the fossil record from Maastrichtian, Late Cretaceous ( Kaim & Beisel 2005).

Etymology. For Takami Nobuhara who made the field trip to the Sada Limestone deposit possible.


University Museum, University of Tokyo