Kaim, Andrzej, Jenkins, Robert G., Tanabe, Kazushige & Kiel, Steffen, 2014, Mollusks from late Mesozoic seep deposits, chiefly in California, Zootaxa 3861 (5), pp. 401-440: 431-432

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Cuspidaria   ? persulcata ( Stanton, 1895)

( Fig. 18 View FIGURE 18 )

Corbula   ? persulcata Stanton, 1895, p. 61–62, pl. 11, fig. 3.

Material and occurrence. One specimen from the Tithonian, Late Jurassic of Paskenta (type locality).

Remarks. Stanton’s holotype ( USNM 23061) is posteriorly more elongate than our specimen but has the same distant concentric furrows which end at the posterior truncation ( Fig. 18 View FIGURE 18 ). Only the external surface of a right valve is available to us, thus a generic placement is difficult. Cuspidariidae   produce very similar shells, including certain species of Cuspidaria Nardo 1840   and Myonera Dall 1886   (i.e., Poutiers & Bernard 1995).

Starting from the careful examination of the Jurassic ‘ Cuspidaria   ’ ibbetsoni, Harper et al. (2003) suggested that many alleged pre-Cretaceous cuspidariids may in fact be corbulids, especially those occurring in brackish environments. Regarding Cuspidaria   ? persulcata from the Paskenta seep locality we suggest the reverse: the species was initially assigned to Corbula   (although hesitantly) and is, in our view, more likely a cuspidariid. Although this classification is based on external shell morphology only, the specimen is from a deep-water environment as are most extant species, and its Tithonian age is just slightly older than that of its sister taxa, the Verticordiidae   and Poromyidae ( Harper et al. 2003)   .


Smithsonian Institution, National Museum of Natural History