Euscelidia artaphernes ( Speiser, 1910 )

Dikow, Torsten, 2003, Revision of the genus Euscelidia Westwood, 1850 (Diptera: Asilidae: Leptogastrinae), African Invertebrates 44 (2), pp. 1-131 : 45-48

Euscelidia artaphernes ( Speiser, 1910) ( E. datis species-group)

Figs 26 , 27A–C, G

Leptogaster artaphernes Speiser, 1910: 83 .

Euscelidia artaphernes, Oldroyd 1939: 28 ; Lindner 1955: 28; Oldroyd 1980: 356.

Redescription: Head: Black; fc yellow pruinose, often long yellow setae originating at antennal base directing ventrally, fc gib distinct; mystax yellow, many macrosetae, nearly as long as prob; oc tr apruinose; occ silver pruinose, setae yellow; Antennae - scp brown, setae yellow; ped light brown, white pruinose, setae yellow and brown; pped light brown proximally, brown distad, white pruinose posteriorly; apsel hyaline; Thorax: Black; sct predominantly black, antero-laterally orange; margins silver or yellow pruinose, yellow setae scattered on surface, setae always directing to the centre; sctl white pruinose, ds sctl s and sctl s long, yellow; Legs - orange to brown; fem light brown, meta fem pale yellow proximally, brown medially, orange distally, clubbed, club densely covered with short, erect yellow setae; pro and meso tib light brown, yellow stripe anteriorly, meta tib orange proximally, brown distad, yellow stripe not reaching tip anteriorly; emp distinct, very short; Wings - densely covered with microtrichia, brown coloured; ptero indistinct, brown; cell d terminating in 2 veins; hlt light brown; Abdomen: Brown to dark brown; T orange on anterior and posterior margins, grey pruinose, setae yellow, T1 with long yellow setae, T 2 in proximal 2/3 with yellow setae laterally; ơ terminalia Figs 27A–C, G - sur bent inwards distally; hypd with membranous area ventrally; d aed shea long, tip Fig. 27G , lat apod bifurcated.

Type material - The ơ lectotype, here designated to preserve stability and make more universal the use of this name, is labelled ‘ Kilimandjaro Sjöstedt. 1905–6 / Kibonoto 1000–1300 m / 29 april (‘29’ handwritten) / Type! Leptogaster artaphernes m P. Speiser det. (handwritten except for ‘P. Speiser det.’) / LECTOTYPE Leptogaster artaphernes

A B C

D E F G H

Speiser, 1910 by T. Dikow 2001 (red label)’. The specimen is directly mounted, is in good condition (sct cracked), and is deposited in the NHRS.

The ^ paralectotype is labelled ‘ Kilimandjaro Sjöstedt. 1905–6 / Kibonoto 1000–1300 m / 4 dec. [4 December] (‘4’ handwritten) / Leptogaster artaphernes m P. Speiser det. (handwritten except for ‘P. Speiser det.’) / PARALECTOTYPE Leptogaster artaphernes Speiser, 1910 by T. Dikow 2001 (yellow label)’. The specimen is directly mounted, is in poor condition (right pped, left meso leg, right meta leg broken; sct cracked; left meta leg attached to the body with glue), and is deposited in the NHRS. Another ^ paralectotype is labelled ‘ Kilimandjaro Sjöstedt. 1905–6 / Obstgarten - Steppe / 22 Mars [March] (‘22’ handwritten) / Leptogaster artaphernes m P. Speiser det. (handwritten except for ‘P. Speiser det.’) / Paratypus (red label with black submarginal border) / 10 52 (red label) / PARALECTOTYPE Leptogaster artaphernes Speiser, 1910 by T. Dikow 2001 (yellow label)’. The specimen is directly mounted, is in poor condition (left pped, left meso leg, and abdomen broken (attached on a piece of cardboard); sct cracked), and is deposited in the NHRS.

Type locality and distribution ( Fig. 26 ): Tanzania, Kilimandjaro area, Kibonoto (= Kibongoto), 310'S 37 05'E. Burundi, Cameroon, DR Congo, Ethiopia, Kenya, Malawi, Mozambique, South Africa, Tanzania, Togo, Uganda, Zambia, Zimbabwe .

Material examined: BURUNDI: 3ơơ Rumonge , 3 58'S 29 26'E, 21.i.1950 ; CAMEROON: 3ơơ Bossoum , 9 56'N 13 41'E, 1.iv., 18.v., 1–10.vi.1914 ; DR CONGO: 1ơ Kivu, 3 03'S 28 06'E; 2^^ Lulua , 5 55'S 25 46'E, iv.1933 ; 1ơ Albertville , 5 55'S 29 12'E, xii.1918 ; 1ơ Baudouinville , 7 03'S 29 47'E, 19.i.1933 ; ETHIOPIA: 2ơơ Harer, 09 19'N 42 07'E; KENYA: 1ơ Lake Victoria , 0 09'S 34 51'E, v.1912 ; 1^ Kuja Valley , 0 48'S 34 34'E, 30.iv.–1.v.1911 ; 2^^ Gelegele River , 0 53'S 35 02'E, vi.1913 ; 1ơ Nairobi , 1 16'S 36 48'E, xi.–xii.1911 ; 1^ Kibwezi , 2 24'S 37 57'E, 2–4.iv.1911 ; MALAWI:1^ Mponela , 13 30'S 33 42'E, 26.ii.1987 ; 1^ Senga Bay , 13 42'S 34 37'E, 7–8.iii.1987 ; 3^^ 1ơ 1? Zomba, 15 18'S 35 10'E; MOZAMBIQUE: 1ơ Nampula, 15 00'S 39 00'E, 19.ii.1982 ; SOUTH AFRICA: Northern Prov: 1ơ Ofcolaco , 2430AB, 20.ii.1980 ; TANZANIA: 1^ Ukerewe, 2 01'S 32 58'E, 1911; 1^ Dar-es-Salam , 6 48'S 39 16'E, 11–20.xii.1961 ; 1^ vii.1902; 1ơ NW Tanganyika, 1910; TOGO: 1ơ Bismarckburg , 811'N 0 41'E, 9–11.v.1893 ; UGANDA: 1^ 2ơơ Abi Farm , 3 05'N 30 56'E, 12.xi.1958 ; 1^ 2ơơ Ankole , 0 43'N 30 11'E, 16.xii.1962, 23–29.iv.1968 ; 1^ 2ơơ Mubende, 0 35'N 31 22'E, 19+ 22.iv.1966 ; 1^ L. Namugabo , 0 04'N 32 15'E, 3.vi.1963 ; 2^^ Bugalla Island , 0 38'S 32 18'E, v.1912 ; 1^ 1ơ 1? Entebbe , 0 03'N 32 27'E, 28.x.1912, 23.v.1913 ; 2ơơ 1? Kome Island , 0 05'S 32 44'E, 15.xii.1918 ; 2^^ Mbarara, 0 36'S 30 38'E, 22–24.x.1911, 15.xi.1934 ; 1^ 1ơ Lolui Island , 0 07'S 33 42'E, v.1965 ; ZAMBIA: 1^ 1ơ Kasanka National Game Reserve , 12 30'S 30 15'E, 16–22.xii.1989 ; 1^ Mungu , 15 16'S 23 04'E, 10.xii.1955 ; ZIMBABWE: 1ơ Harare, 17 46'S 31 02'E, i.1969 ; 1ơ Bindura , 17 18'S 31 19'E, i.1929 ; COUNTRY UNKNOWN: 1^ Kawanda [Malawi, Uganda, or Zambia], 9.xi.1957 ; 1ơ Abyssinia , xi.1911 ; 1ơ no locality data. Depository : BMNH, ISNB, MRAC, NHMW, NMSA, SMNS, USNM, ZMHB, ZSMC .

Remarks: Illustration of ơ terminalia by Theodor (1976: 20) is not artaphernes but datis .

Figures

Fig. 26. Distribution. E. artaphernes, circles; E. datis, squares; E. lata sp. n., triangles; E. milva sp. n., stars; ^^ of species-group, plus sign.

Fig. 27. ơ terminalia. A–C, G. Euscelidia artaphernes. A. Dorsal. B. Ventral. C. Lateral. G. Aedeagus lateral. D–F, H. E. rapax. D. Dorsal. E. Ventral. F. Lateral. H. Aedeagus lateral (tip broken and shape unknown; aed apod not illustrated). Scale line = 1 mm.