Euscelidia nenemusha ( Speiser, 1910 ) Dikow, 2003

Dikow, Torsten, 2003, Revision of the genus Euscelidia Westwood, 1850 (Diptera: Asilidae: Leptogastrinae), African Invertebrates 44 (2), pp. 1-131 : 85-88

Euscelidia nenemusha ( Speiser, 1910) , comb. n.

Fig. 50

Leptogaster nenemusha Speiser, 1910: 84 ; Hull 1962: 300; Oldroyd 1980: 358.

Diagnosis: The species is distinguished, together with castanea and schoutedeni , from congeners by the overall brown colour, the predominantly apruinose sct, and the apruinose T 8 in the ^^.

Redescription: Head: Black; fc silver pruinose, fc gib indistinct, mystax white, 11 macrosetae; oc tr white pruinose dorsally; occ yellow pruinose, few white setae; Antennae - scp light brown, setae light brown; ped and pped yellow, white pruinose; apsel light brown; Thorax: Brown; sct brown, apruinose, anterior, lateral, and posterior margins yellow pruinose, 2 black spots laterally, few yellow setae on surface, macrosetae: black, 1 npl s, 1 spal s; sctl silver pruinose, ds sctl s and sctl short, yellow; Legs - yellow to light brown; pro and meso fem brown anteriorly, yellow posteriorly, meta fem pale yellow proximally, brown dorsally, yellow ventrally, slightly clubbed, white setae; tib light brown, yellow stripe not reaching tip anteriorly; emp about a quarter of length of clw; Wings - hyaline, some microtrichia distally; ptero distinct, brown; cell d terminating in 2 veins; hlt yellow with brown knob; Abdomen: Brown, T white pruinose, T1 with black setae, S grey pruinose; ơ terminalia - ơơ specimens unknown.

Type material - The ^ holotype is labelled ‘ Meru 3,000 –3,500 m (3,500’ crossed out) / Sjöstedt. / jan. / Typus (red label with black submarginal border) / 11 52 (red label) / Type! Leptogaster nenemusha m P. Speiser det. (handwritten except for ‘P. Speiser det.’) / HOLOTYPE Leptogaster nenemusha Speiser, 1910 by T. Dikow 2001 (red label)’. The specimen is directly mounted, is in good condition, and is deposited in the NHRS.

Type locality and distribution ( Fig. 50 ): Tanzania , Meru, 3 13'S 36 44'E. Tanzania .

Remarks: The species is only known from the female holotype. It is related to castanea and schoutedeni , which are also only known from females. It is necessary to examine male specimens from the type localities of all three species to discuss a probable synonymy.

Euscelidia nitida (Wiedemann, 1828)

Figs 51A–D , 53

Leptogaster nitidus Wiedemann, 1828: 535 ; Loew 1858: 352; 1860: 102; Hull 1962: 300.

Euscelidia nitida, Engel 1929: 154 ; Oldroyd 1974: 20; Lehr 1988: 266.

Diagnosis: The species is distinguished from congeners by the densely grey pruinose sct with two narrow median apruinose stripes and two lateral apruinose stripes, the apruinose proximal half of abdominal T2, and features of the ơ terminalia.

Redescription: Head: Black; fc silver pruinose, fc gib distinct, mystax white, few macrosetae; oc tr grey pruinose dorsally; occ grey pruinose, setae white; Antennae - scp light brown, setae brown; ped light brown, setae brown; pped light brown, white pruinose dorsally in proximal half of length; apsel brown; Thorax: Black; sct grey pruinose, 2 narrow median stripes, not reaching posterior margin, and 2 lateral stripes, not reaching anterior and posterior margins, apruinose; sctl grey pruinose, ds sctl s and sctl very short, white; Legs - yellow to brown; pro and meso fem yellow, meta fem pale yellow proximally, clubbed, club brown; tib light brown, pale yellow stripe not reaching tip on meta tib anteriorly, meta tib brown in distal half; emp about a quarter of length of clw; Wings - hyaline, very few microtrichia on remigium; ptero distinct, brown; cell d terminating in 3 veins; hlt yellow; Abdomen: Black; T predominantly brown pruinose, anterior, lateral, and posterior margins grey pruinose, T 2 in proximal half with apruinose spot, T1 with white setae, T2 with few white setae laterally, S grey pruinose; ơ terminalia Figs 51A–D - sur pointed distally; lobe ventrally; hypd ventral margin straight, distally with 2 pointed lobes; d aed shea tubular; lat apod bifurcated.

Type material - The ^ lectotype, here designated to preserve stability and make more universal the use of this name, is labelled ‘Nubien Rüppell (handwritten) / L. nitidus Wied. * (handwritten) / (small red label) / 9973 / Holotypus (red label) / LECTOTYPE Leptogaster nitidus Wiedemann, 1828 by T. Dikow 2000 (red label)’. The specimen is directly mounted, is in fairly poor condition (left meso and meta leg, right meta leg, and left wing broken), and is deposited in the ZMHB.

The ^ paralectotype is labelled ‘ Abyssinia Dr. Rüppell. / 113 (blue label with black submarginal border) / Leptogaster nitidus Wied. (handwritten) / Typus (red label with black submarginal border) Dipt. 87 (on white backside of label) / PARALECTOTYPE Leptogaster nitidus Wiedemann, 1828 by T. Dikow 2000 (yellow label)’. The specimen

A B C D E

G H I is in poor condition (right antenna, left meso and meta legs broken, specimen greasy) and is deposited in the SMFD. The paralectotype of undeterminable gender is labelled ‘ Abyssinia Dr. Rüppell. / Paratypoid (red label with black submarginal border) Dipt. 87a (on white backside of label) / PARALECTOTYPE Leptogaster nitidus Wiedemann, 1828 by T. Dikow 2000 (yellow label)’. The specimen is directly mounted, is in poor condition (antennae, left pro tarsus, left meso leg, right meta leg, and abdomen broken), and is deposited in the SMFD .

Type locality and distribution ( Fig. 53 ): Nubien – Nubia is a classic term for an ancient kingdom in north-eastern Africa , an area today probably comprising Egypt and Sudan, no co-ordinates available. Probably Egypt and Sudan. Additionally to the specimens examined the species is known to occur in Arabia ( Lehr 1988; Oldroyd 1980) .

Material examined: EGYPT: 1ơ no exact locality data. Depository: NHMW.

Figures

Fig. 50. Distribution. Euscelidia castanea, circles; E. hyalina sp. n., triangle; E. nenemusha, star; E. obudu sp. n., pentagon; E. schoutedeni, squares.

Fig. 51. ơ terminalia. A–D. Euscelidia nitida. A. Dorsal. B. Ventral. C. Lateral. D. Aedeagus lateral. E–I. E. obtusa sp. n. E. Aedeagus lateral. F. Aedeagus dorsal. G. Dorsal. H. Ventral. I. Lateral. Scale line = 1 mm.

Fig. 53. Distribution. Euscelidia nitida, triangle; E. pallasii, squares.