Glyphiulus deharvengi, Golovatch & Geoffroy & Mauriès & Spiegel, 2007

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Spiegel, Didier Van Den, 2007, Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group, Zoosystema 29 (1), pp. 7-49 : 32-34

publication ID	https://doi.org/ 10.5281/zenodo.5392079
persistent identifier	https://treatment.plazi.org/id/03F4C378-A236-3053-FC80-0210FE2FF935
treatment provided by	Marcus
scientific name	Glyphiulus deharvengi
status	sp. nov.

Treatment

Glyphiulus deharvengi View in CoL n. sp.

( Figs 19 View FIG ; 20 View FIG )

TYPE MATERIAL. — China. Hunan Prov., Longshan: Huoyan, Feihu Dong Cave ( Flying Tiger Cave ), 11.VIII.1995, leg. A. Bedos & L. Deharveng (CHI- 022), holotype ♂ ( MNHN GA 040 View Materials ) ; paratype 1 ♂ ( MNHN GA 040 View Materials ). — Hunan Prov., Longshan, Huoyan, Baiyan Dong Cave ( White Rocks Cave ), 15.VIII.1995, leg. A. Bedos & L. Deharveng (CHI- 032), paratypes 1 ♂, 1 ♀, 1 ♀ juv. ( MNHN GA 040 View Materials ) ; 1 ♂ ( ZMUM). — Hunan Prov., Longshan: Huoyan, Renmi Dong Cave , 19.VIII.1995, leg. A. Bedos & L. Deharveng (CHI-043) ; paratypes 1 ♂ ( MNHN GA 040 View Materials ) ; 1 ♀ ( SEM) .

ETYMOLOGY. — Honours Louis Deharveng, one of the collectors.

DIAGNOSIS. — Differs by the relatively large size, the nearly completely smooth collum, the complete depigmentation (even of ocellaria), the elongated legs (with claws devoid of a spine at base from leg 4 onwards), the low, but evident, transverse pleural ridge behind the gonopod opening on male segment 7, the bare but flagelliform distal process of the posterior gonopods, etc.

DESCRIPTION

Length of adults of both sexes 40-55 mm, width 2.0- 2.3 mm. Coloration usually uniformly dark brown due to heavy sclerotization, tegument being coloured only due to chitin; sometimes ozoporiferous tubercles reddish.

Adults with 55-68p+4-1a+T; juvenile ♀ with 40p+5a+T, recognised by considerably reduced ozoporiferous tubercles on segments 7-33, plus 5a+T. Length of holotype about 55 mm, width 2.3 mm, with 67p+1a+T.

Antennae slightly more elongate than usual ( Figs 19A View FIG ; 20A View FIG ). Ocellaria unpigmented, barely traceable, with about 10 or 11 ocelli in largest specimens. Gnathochilarium ( Fig. 20B View FIG ) with an undivided mentum (n = 1).

Carinotaxy of collum unusual in that all crests are nearly obliterated ( Fig. 19A, B View FIG ), though a typical pattern is traceable. Carinotaxy of subsequent metaterga typical ( Fig. 19 View FIG A-C, E), but crests low, especially dorsal ones on segments 2-6; only two lateral crests below ozoporiferous ones, divided from segment 7 onwards, followed by a dozen striae. A low, but evident, transverse pleural ridge behind gonopod opening on male segment 7. Epiproct slightly elongate apically, more narrowly rounded than usual; hypoproct more narrowly emarginate at caudal margin ( Fig. 19D View FIG ).

Legs elongate ( Fig. 20F View FIG ), claws strongly elongate, without a spine near base starting from leg 4 ( Fig. 19F View FIG ).

Male legs 1 with vestigial tubercular coxites ( Fig. 20C View FIG ). Penes with 4 or 5 long setae distolaterally ( Fig. 20D View FIG ), male legs 3 elongate, much as in G. granulatus ( Fig. 20E View FIG ). Anterior gonopods with small subunciform protuberances at median corners of coxosternum ( Fig. 20G View FIG ). Posterior gonopods with a flagelliform but bare distal process ( Fig. 20H View FIG ).