Cheilolejeunea trifaria (Reinw, Blume & Nees) Mizut.

Silva, Jainara Pereira, Oliveira-da-Silva, Fúvio Rubens, Ilkiu-Borges, Anna Luiza & Fernandes, Rozijane Santos, 2021, Leafy liverworts of Chapada das Mesas National Park: a floristic survey and checklist of the leafy liverworts of Maranhão state, Brazil, Check List 17 (1), pp. 479-495 : 486

** Cheilolejeunea trifaria (Reinw, Blume & Nees) Mizut.

Figure 3H

Material examined. BRAZIL • Maranhão, município de Carolina, Parque Nacional Chapada das Mesas, Riacho Buenos Aires, corticícola, ilha da Cachoeira do

Prata; 06°56′S, 047°20′W; alt. 228 m; 30 Oct. 2017; JAS Silva 149; CCAA 1567.

Identification. Plants olive-green. Ventral merophyte of 2 cells wide. Leaves subimbricate, suborbicular, apex rounded, plane to slightly incurved, margin entire or sinuate, dorsal and ventral margin curved; median cells isodiametric, ca. 20‒25 µm, trigones small to large. Lobules ovate to elongate (usually hidden behind the underleaves), 1/3‒1/2 lobe length, apical tooth short. Underleaves contiguous to subimbricate, bifid to 1/3‒1/2 of length, ovate, larger than long, 3‒6 × stem width, base broadly rounded with insertion line deeply arched, apex acute, margin entire. Gynoecia with 1‒2 lejeuneoid innovation, often fertile. Perianth obovate, 5-keeled.

Cheilolejeunea trifaria is easily recognized by relatively large plants with suborbicular leaf lobes, lobules ovate to elongate, hidden behind the large underleaves (3‒6 × stem width).

Description and illustration. Gradstein and Ilkiu- Borges (2009: 68, fig. 38e–g).

Figures

Figure 3. Leafy liverworts of the Chapada das Mesas National Park; A. Frullania intumescens; B. Frullania platicalyx; C. Frullania rio-janeirensis; D. Ceratolejeunea cubensis; E. Cheilolejeunea acutangula; F. Cheilolejeunea adnata var.adnata; G. Cheilolejeunea intertexta; H. Cheilolejeunea trifaria; I. Lejeunea juruana; J. Lejeunea quinqueumbonata; K. Metalejeunea cucullata; L. Zoopsidella integrifolia; M. Zoopsidella macella; N. Lophocolea bidentata; O. Plagiochila raddiana; P. Plagiochila simplex (Scale bar: A-B, F-G, P= 20 µm; C-D, H-O= 100 µm; E= 300 µm).