Clypeocaenis

Vannachak, Viengkhone, Malzacher, Peter & Sangpradub, Narumon, 2024, Contribution to the genus Clypeocaenis Soldán, 1978 (Ephemeroptera: Caenidae) from Lao People’s Democratic Republic with a key to Oriental species, Zootaxa 5501 (4), pp. 501-523 : 517-518

publication ID

https://doi.org/ 10.11646/zootaxa.5501.4.1

publication LSID

lsid:zoobank.org:pub:29E9284E-6F5C-46E5-BA25-B54F820BE0C0

DOI

https://doi.org/10.5281/zenodo.13753966

persistent identifier

https://treatment.plazi.org/id/03F55817-FFBD-FFDE-FF6D-FA56FA2DFC27

treatment provided by

Plazi

scientific name

Clypeocaenis
status

 

Key to the larvae of Oriental Clypeocaenis View in CoL View at ENA species

1 Clypeal protrusion with 4 or more setae.................................................................... 2

– Clypeal protrusion with two setae........................................................................ 4

2 Clypeal protrusions with 20–35 long setae.These setae, along with those on the body surface, often possess many thin, irregularly arranged, and matted branches, creating what are referred to as sausage-shaped setae ( Malzacher 2013, Figs. 37–39).................................................................................................. Cl. soldani

– Clypeal protrusion with less than 20 long setae. Surface without sausage-shaped bristles, but with shield-shaped microtrichia ( Fig. 9A, C View FIGURE 9 ).......................................................................................... 3

3 Clypeal protrusions with 10–15 long setae ( Fig. 9A View FIGURE 9 ). The cuticle features shield-shaped, spider web-like microtrichia with smooth margins ( Fig. 8C View FIGURE 8 ).................................................................... Cl. multisetosa View in CoL

– Clypeal protrusion with about 4 long setae and several short ones ( Fig. 6A View FIGURE 6 ). Shield-shaped microtrichia with frayed margin ( Fig. 8A View FIGURE 8 )................................................................................. Cl. oligosetosa View in CoL

4 Mid and hind femora with long, acute marginal setae ( Soldán 1978, Figs. 9 View FIGURE 9 , 10 View FIGURE 10 , 21). Operculate gill’s length more than twice its width, with blunt or spatulate marginal setae ( Balasubramanian & Muthukatturaja 2021, Fig. 31).................... 5

– Mid and hind femora with short marginal setae ( Fig. 3D, E View FIGURE 3 ). Operculate gill with very short length not more than 2 time as width, blunt or spatulate marginal setae, on lateral margin very short or absent ( Balasubramanian & Muthukatturaja 2021, Fig.14 View FIGURE 14 )............................................................................................. 8

5 Fore tarsus and the distance between the transverse rows of bases of filtering setae on the foretibia are of equal length ( Soldán & Landa 1991, Fig. 14 View FIGURE 14 )................................................................... Cl. femorisetosa View in CoL

– Fore tarsus 1.7 to 2 times as long as the distance between the transverse rows of bases of filtering setae on the foretibia ( Soldán 1983, Fig. 12 View FIGURE 12 )........................................................................................ 6

6 Genae margin angled. Foreleg with feathered bristles in some places, e.g. marginal on coxae and femora ( Muthukatturaja et al. 2020, Fig. 27).............................................................................. Cl. sharadhae

– Genae differently shaped. Fore leg without feathered bristles................................................... 7

7 Apical margins of the clypeal protrusion straight ( Soldán 1978, Fig. 16). The third segment of the labial palp hardly widened basally ( Soldán 1978, Fig. 3 View FIGURE 3 ).................................................................... Cl. bisetosa View in CoL

– Apical margin of the clypeal protrusion rounded. The third segment of the labial palp significantly widened basally ( Balasubramanian & Muthukatturaja 2021, Figs. 2 View FIGURE 2 , 26).............................................. Cl. napoklu

8 Clypeal protrusions markedly large, apically domed. Genae protruding, doubly angled, the margin between angles straight ( Srinivasan et al. 2022, Fig. 2A View FIGURE 2 ).............................................................. Cl. malzacheri

– Clypeal protrusion flatly rounded, not domed apically........................................................ 9

9 Setae of transverse row on forefemur broad ( Fig. 3A, B View FIGURE 3 ). Segment I of maxillary palp narrow, about half as wide as galea-lacinia ( Fig. 2D View FIGURE 2 ). Setae on outer margin of mid and hind femora spatulate, with a very thin basal part ( Fig. 3E, G View FIGURE 3 ). Claws of mid and hind legs with 3–5 denticles ( Fig. 3F, H View FIGURE 3 ). Ventral side of operculate gill without microtrichia ( Figs. 4G View FIGURE 4 , 12F View FIGURE 12 )............................................................................................. Cl. laosensis sp.nov.

– Setae of transverse row on forefemur narrow. Segment I of maxillary palp broadened, about as wide as galea-lacinia. Setae on outer margin of mid and hind femora thin, acute or bifid. Claws of mid and hind legs with 8–10 denticles ( Balasubramanian & Muthukatturaja 2021, Figs. 7 View FIGURE 7 , 10–12 View FIGURE 10 View FIGURE 11 View FIGURE 12 ;). Ventral side of operculate gill with a sublateral band of microtrichia ( Fig. 10D View FIGURE 10 ).... 10

10 Pedicel two and a half time as long as third antennal segment; the latter about as long as the following segments; genae flatly angled. Segment III of labial palp basally strongly broadened Coxae enlarged, two third as wide as length of fore femur. Median ridge of operculate gill scarcely keeled ( Muthukatturaja et al. 2020, Figs. 1 View FIGURE 1 , 7–10 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 , 14 View FIGURE 14 )...................... Cl. gayathri

– Pedicel one and a half times as long as the third antennal segment; the latter distinctly longer than the following segments; genae margin not angled. Segment III of the labial palps not strongly broadened basally ( Balasubramanian & Muthukatturaja 2021, Fig. 9 View FIGURE 9 ). Coxa half as wide as the length of the forefemur. The median ridge of the operculate gill strongly keeled ( Balasubramanian & Muthukatturaja 2021, Figs. 1 View FIGURE 1 , 10–12 View FIGURE 10 View FIGURE 11 View FIGURE 12 , 14 View FIGURE 14 ).......................................... Cl. kaveri

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Ephemeroptera

Family

Caenidae

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