Coelotes thailandensis Dankittipakul & Wang 2003

Wang, Xin-Ping, Griswold, Charles E. & Ubick, Darrell, 2009, On the pseudoterrestris species group of the spider genus Coelotes (Araneae, Amaurobiidae), Zootaxa 2313, pp. 1-34 : 26-28

Coelotes thailandensis Dankittipakul & Wang 2003

Figs 128–142 ; 157

Coelotes thailandensis Dankittipakul & Wang 2003: 735 , figs 24–25 (male holotype, in MHNG, examined). – Dankittipakul, Chami-Kranon & Wang 2005: 7 , figs 9-12.

Material examined: Holotype. THAILAND: Chiang Rai: 3, Wiang Pa Phao District, Khun Jae National Park, N19°05'32", E99°50'41", Doi Mae Toh, Doi Mae Hoh Waterfall, October 15, 2002, P. Sankittipakul & S. Sonthichai ( MHNG).

Other material examined. THAILAND: Chiang Mai: 1Ƥ, Chom Thong District, Doi Inthanon National Park, Doi Inthanon, N18°35'14", E98°29'7", evergreen hill forest, 1530–1680 m, December 21, 2004, T. Chami-Kranon & S. Sonthichai ( MHNG).

Diagnosis: This species resembles C. pseudoterrestris in having the short, laterally arising epigynal teeth, similar atrium, anteriorly extending copulatory ducts and similar palp, but can be distinguished by the short, round copulatory ducts (less than spermathecal length), blunt distal end of patellar apophysis, and subequal size of conductor and its dorsal apophysis ( Figs 128–131 ).

Description: Male (holotype): Large sized coelotine, total length 11.67 ( Fig. 137 ). AME smallest, ALE largest, posterior eyes subequal in size, slightly larger than AME; anterior eyes equally separated by 2/3 of AME diameter; PME separated from each other by ¾ of their diameter, from PLE by about 1.5 times PME diameter ( Fig. 138 ). Promargin of chelicera with 3 teeth, retromargin 4. Labium with length and width subequal (L/W=1.03) ( Fig. 139 ). Palpal patellar apophysis large, without pointed distal end; RTA more than half of tibial length, extends distally; lateral tibial apophysis short, broad, distinctly separated from RTA; cymbial furrow short, about 1/3 of cymbial length; conductor short, about the same size as its dorsal apophysis, with a small basal lamella; median apophysis broad, round, spoon-shaped; embolus short, filiform, prolateral in origin, with thread start from middle part of embolic base ( Figs 130–131 , 134–136 ).

Female: Large sized coelotine, total length 10.63 ( Fig. 140 ). AME smallest, lateral eyes subequal in size, PME slightly larger than AME; AME separated from each other by ¾ of their diameter, from ALE by about 1/ 3 of AME diameter; PME separated from each other by slightly less than their diameter, from PLE by slightly more than PME diameter ( Fig. 141 ). Promargin of chelicera with 3 teeth, retromargin 4. Labium longer than wide (L/W=1.20) ( Fig. 142 ). Epigynal teeth short, arising laterally at level between anterior atrial margin and epigastric furrow, separated by more than atrial width; atrium small, with distinct, continuous anterior margin but indistinct lateral margins; copulatory ducts large, contiguous, extending anteriorly, more or less round; spermathecae broad, closely set or slightly separated, slightly extending anteriorly; spermathecal heads short, indistinct, arising anteriorly ( Figs 128–129 , 132–133 ).

Notes: The female specimen was identified by Dankittipakul et al. (2005) as C. thailandensis but shows a very different labium size ratio than that of the male holotype. The male holotype, from Khun Jae National Park, Chiang Rai Province, is subequal in labium length and width (L/W=1.03), while the female, from Doi Inthanon National Park, Chiang Mai Province which is about 100 km away from locality of the holotype male ( Dankittipakul et al. 2005), is much longer than wide (L/W=1.20). The male labium usually has a larger ratio of length and width than the female. In addition, the male specimens from Doi Inthanon National Park also showed a much shorter patellar apophysis (less than half of patellar length) than those from Khun Jae National Park (as long as patellar length) ( Dankittipakul et al. 2005: figures 11, 12). Further collection of both sexes from both localities is needed to adequately demonstrate their species status.

Distribution: Thailand (Chiang Mai, Chiang Rai) ( Fig. 157 ).

Figures

FIGURES 157. Distribution records of the pseudoterrestris group species: Coelotes suthepicus and C. thailandensis.

FIGURES 128 – 131. Coelotes thailandensis Dankittipakul & Wang 2003, male holotype (MHNG) from Khun Jae National Park, Chiang Rai, Thailand and female (MHNG) from Doi Inthanon National Park, Chiang Mai, Thailand. 128,129 Female epigynum, ventral and dorsal view; 130,131 Male palp, ventral and retrolateral view.

FIGURES 132 – 136. Coelotes thailandensis Dankittipakul & Wang 2003, male holotype (MHNG) from Khun Jae National Park, Chiang Rai, Thailand and female (MHNG) from Doi Inthanon National Park, Chiang Mai, Thailand. 132,133 Female epigynum, ventral and dorsal view; 134 – 136 Male palp, prolateral, ventral and retrolateral view.

FIGURES 137 – 142. Coelotes thailandensis Dankittipakul & Wang 2003, male holotype (137 – 139) from Khun Jae National Park, Chiang Rai, Thailand (MHNG) and female (140 – 168142 from Doi Inthanon National Park, Chiang Mai, Thailand (MHNG). 137,140 Habitus, dorsal view; 138,141 Eyes, view between front and dorsal; 139,142 Labia, ventral view.