Zischkaia amalda ( Weymer, 1911 ), Weymer, 1911

Nakahara, Shinichi, Zacca, Thamara, Dias, Fernando M. S., Dolibaina, Diego R., Xiao, Lei, Espeland, Marianne, Casagrande, Mirna M., Mielke, Olaf H. H., Lamas, Gerardo, Huertas, Blanca, Kleckner, Kaylin & Willmott, Keith R., 2019, Revision of the poorly known Neotropical butterfly genus Zischkaia Forster, 1964 (Lepidoptera, Nymphalidae, Satyrinae), with descriptions of nine new species, European Journal of Taxonomy 551, pp. 1-67: 20-22

publication ID

https://doi.org/10.5852/ejt.2019.551

publication LSID

lsid:zoobank.org:pub:C3C851C3-0F12-412C-A15B-56F0F263CD00

DOI

http://doi.org/10.5281/zenodo.3477358

persistent identifier

http://treatment.plazi.org/id/03F587B3-FF89-FF9A-0702-F9A91561FDD7

treatment provided by

Plazi

scientific name

Zischkaia amalda ( Weymer, 1911 )
status

 

Zischkaia amalda ( Weymer, 1911)  

Figs 2 View Fig A–B, 4 View Fig A, 14 View Fig

Euptychia amalda Weymer, 1911: 213   , pl. 48, fig. f[2] (Type Locality: Mapiri [La Paz, Bolivia]).

Euptychia amalda – Gaede 1931: 437   . — D’Abrera 1988: 789.

Zischkaia amalda – Forster 1964: 116–117   , fig. 124. — Lamas 2004: 223.

Material examined

Lectotype (here designated)

BOLIVIA • ♂; “//CoTypus Nr. Euptychia amalda (Stgr)Weymer Zoologische Staatssammlung München.   // amalda Weym   [er].// Mapiri [La Paz, Bolivia]// 521.// 261.// Original [i.e., Type]// Genitalic vial SN-17-21 S. Nakahara //”; ZSM.  

Other material (12 ♂♂)

BOLIVIA – La Paz • 1 ♂; Coroico ; [16°10′ S, 67°44′ W]; 1200 m; Fassl A.H. leg.; ZSM GoogleMaps   1 ♂; Río Zongo ; [16°3′40″ S, 68°1′2″ W]; 1200 m; 1895–1896; Garlepp leg.; MNHU GoogleMaps   1 ♂; Zongo [= Río Zongo]; [16°7′ S, 68°2′ W]; Garlepp leg.; MNHU GoogleMaps   1 ♂; same collection data as for preceding; BMNH(E)-10247952; NHMUK GoogleMaps   .

PERU – Huánuco • 1 ♂; Tingo María ; [9°18′ S, 76°0′ W]; 800 m; 7 Aug. 1979; FLMNH- MGCL-1036040; FLMNH GoogleMaps   . – Junín • 1 ♂; Chanchamayo ; [11°4′ S, 75°19′ W]; 1000–1400 m; Jun.– Aug. 1901; W. Hoffmanns leg.; BMNH(E)-1718080; NHMUK. – Not located GoogleMaps   6 ♂♂; “Peru” [most likely from Bolivia]; USNM   .

Identification and taxonomy

The male of Zischkaia amalda   is distinguished from Z. baku   sp. nov. and Z. chullachaki   sp. nov. by the VFW submarginal line not undulating (the VFW submarginal line is undulating in Z. baku   sp. nov. and Z. chullachaki   sp. nov.), with the area of dorsal androconia rather reduced and the androconia not as prominent as in those two species. Furthermore, the male genitalia can be used to distinguish this species from Z. baku   sp. nov., with the winglet of the aedeagus being developed ( Fig. 4A View Fig ) (reduced in Z. baku   sp. nov.), and the anterior margin of the tegumen being concave in dorsal view in Z. amalda   , whereas it is rather straight or convex in Z. baku   sp. nov. and Z. chullachaki   sp. nov. The male of Z. amalda   is distinguished from that of Z. pacarus   by the dorsal androconia being more prominent (dorsal androconia somewhat inconspicuous in Z. pacarus   ), in addition to its smaller adult size ( Z. amalda   : FW length 22– 23 mm (n = 3); Z. pacarus   : 25 mm (n = 5)). The male genitalia of Z. amalda   is distinguished from that of Z. pacarus   by having a subtriangular apical process of the valva (apical process of valva somewhat rectangular with a convex distal margin in Z. pacarus   ). Zischkaia amalda   is perhaps most similar to Z. arctoa   sp. nov., but see the relevant section of Z. arctoa   sp. nov. for diagnostic characters to separate these two species. Also, see the relevant section of Z. arenisca   sp. nov. for diagnostic characters to distinguish Z. amalda   from Z. arenisca   sp. nov. and Z. chullachaki   sp. nov.

Euptychia amalda   was described by Weymer (1911) based on an unstated number of specimens from Mapiri, La Paz, Bolivia. The original description is accompanied by a ventral surface illustration ( Weymer 1911: pl. 48, fig. f[2]), but the description and illustration alone cannot be used to confidently identify this species due to the fact that wing patterns of species of Zischkaia   are often similar. The syntype male, located in the ZSM, has androconia on the DFW and DHW, which is implied in the original description (“The basal and median areas of the forewing above are velvety dark brown, the costal and distal margins lighter brown”), indicating that the description did indeed apply to a species in the “ pacarus   clade” of Zischkaia   . Forster (1964), examined the abovementioned syntype, in addition to another male specimen from Coroico, La Paz, Bolivia (collected by A.H. Fassl), and stated that “this species is apparently widespread in the Yungas, this rare species is similar to the common southern Brazil [Z.] fumata   [i.e., Z. pacarus   ] and it is so similar that it can be seen as subspecies of a single species. In genital apparatus no differences can be observed, however, [Z.] amalda   is smaller with lighter brown color of the dorsal side of the wings so that the androconial spot of the males emerges more clearly”. Forster (1964: 114, fig. 124) correctly identified Z. pacarus   (referred to as Z. fumata   ), but we disagree with his claim of a lack of genitalic differences between Z. amalda   and Z. pacarus   , having observed slight differences between the male genitalia of these two species (see Fig. 4A, G View Fig ). Nevertheless, the specimens of Z. amalda   deposited at the ZSM examined by Forster were not dissected and the origin of the specimen he dissected is unknown. It is therefore not possible to verify whether he dissected Z. amalda   or some other similar species, such as Z. chullachaki   sp. nov., which has similar genitalia. Due to resemblance of this taxon to congeners described in this study, we designate as lectotype of Euptychia amalda Weymer 1911   the male syntype, illustrated by Warren et al. (2018) and deposited at the ZSM, in order to settle the nomenclature (lectotype designation).

Variation

The curvature of the VHW median line below M 3 is apparently variable; the curvature of the phallobase is also apparently variable.

Distribution ( Fig. 14 View Fig )

This species is currently known from the foothills of the eastern Andes, from Tingo María, Huánuco, Peru to Coroico, La Paz, Bolivia.

ZSM

Bavarian State Collection of Zoology

NHMUK

Natural History Museum, London

FLMNH

Florida Museum of Natural History

USNM

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Lepidoptera

Family

Nymphalidae

Genus

Zischkaia

Loc

Zischkaia amalda ( Weymer, 1911 )

Nakahara, Shinichi, Zacca, Thamara, Dias, Fernando M. S., Dolibaina, Diego R., Xiao, Lei, Espeland, Marianne, Casagrande, Mirna M., Mielke, Olaf H. H., Lamas, Gerardo, Huertas, Blanca, Kleckner, Kaylin & Willmott, Keith R. 2019
2019
Loc

Zischkaia amalda – Forster 1964: 116–117

Lamas G. 2004: 223
2004
Loc

Euptychia amalda –

D'Abrera B. 1988: 789
Gaede M. 1931: 437
1931
Loc

Euptychia amalda

Weymer G. 1911: 213
1911