Amboakis diffusa Opitz, 2010

Opitz, Weston, 2010, New taxa of Epiphloeinae Kuwert (Cleridae) and Chaetosomatidae Crowson (Coleoptera: Cleroidea), Insecta Mundi 2010 (123), pp. 1-28 : 6-7

Amboakis diffusa Opitz , new species

Figure 10 , 29 , 49 .

Holotype. Male. ECUADOR, TUNGURAHUA, 38 KM E BAÑOS, 4000' SEP 23 1996 E. GIESBERT, COLL ( FSCA). (Specimen pin mounted, antenna and gender label affixed to paper support card; locality label; acronymic label; Holotype label; plastic vial with abdomen and aedeagus.)

Paratypes. None.

Diagnosis ( Fig. 49 ): From members of Amboakis stenosis Opitz , that also have the posterior region of the elytra flared, the members of this species are distinguished by the much greater expansion of the funicular antennomeres (compare Figure 3, 10 ).

Description. Size: Length 5.0 mm: width 1.4 mm. Integument: Cranium bicolorous, lower frons and postgena dark brown, clypeus, upper frons, and vertex yellow; pronotum dark brown; elytra dark brown; legs bicolorous, profemur mostly dark brown, mesofemur and metafemur mostly yellow, tibiae and tarsi dark brown. Head: Wider than pronotum (28:24); vertex narrower than eye (8:10); funicular antennomeres considerably expanded ( Fig. 10 ). Thorax: Pronotum, quadrate (24:24), side margins more convex than tuberculate, punctations large rendering disc subrugose, discal trichobothrium set in deep, transverse shallow, and spheroid depression; elytra, form narrow triangular, ratio of elytral length to elytral width 3:1, ratio of elytral length to pronotal length 5:1, punctations intermediate in size, seriate and arranged into 10 rows, deeply impressed, punctations wider than width of interstitial spaces, distal margin of epipleuron margin not spinous, disc vested profusely with dark setae; protibial anterior margin with 7 spines. Abdomen: Aedeagus as in figure 29.

Variation. One specimen examined.

Natural History. The Holotype was collected during September at 1220 m.

Distribution. Known only from the type locality.

Etymology. The specific epithet diffusa is a Latin adjective that stems from diffusus (= spread out). I refer to the posterior expansion of the elytra.

Taxonomic Notes. This species belongs in the stenosis species group as defined in Opitz (2006: 144), but the elytral punctations are slightly larger as one finds in specimens of A. stenosis . In Opitz (2006) the included key to Amboakis should be modified to read:

2'. Epipleural and sutural margin not parallel, epipleural margin flared in posterior half ............ 3

3(2'), Funicular antennomeres moderately expanded ( Fig. 3 ) ( México: Oaxaca) .................................... ......................................................................................................... Amboakis stenosis Opitz

– Funicular antennomeres extensively expanded ( Fig. 10 ) ( Bolivia: Tungurahua) .......................... ......................................................................................................... Amboakis diffusa Opitz

Amboakis demagna Opitz , new species

Figure 12 , 27 , 50 .

Holotype. Male. ECUADOR: Orellana: Res. Ethnica Waorani, 1 km S Onkone Gare Camp , 00 o 39’S 76 o 26’W, 22-VI- 1996, 216 m, Fogging terre firme T.L. Erwin ( USNM). (Specimen point mounted, gender label affixed to paper point; locality label; second Natural History label reads Fogging terre firme; USNM acronymic label; Holotype label.)

Paratypes. None.

Diagnosis ( Fig. 50 ): Within Amboakis , the members of this species resemble most those of A. prolata Opitz , from which they are conveniently separated by the brief pale line that extends backwards from the elytral humeral margin.

Description. Size: Length 4.0 mm: width 1.0 mm. Integument: Predominantly dark brown; mouthparts, antenna, and legs yellow; elytra mostly dark brown, with short yellow line extending backwards from humeral margin and yellow transverse band at middle. Head: Wider than pronotum (28:23); vertex narrower than eye (8:11); funicular antennomeres slightly expanded, last club antennomere very large ( Fig. 12 ). Thorax: Pronotum, transverse (55:45), side margins behind anterior transverse depression more convex than tuberculate, punctations large rendering surface subrugose, discal trichobothrium set in deep, transverse shallow, and spheroid depression; elytra, form short rectangular, ratio of elytral length to elytral width 2:4, ratio of elytral length to pronotal length 4:1, punctations large, seriate, shallowly impressed, wider than width of interstitial spaces, distal margin of epipleuron margin minutely spinous, disc vested profusely with dark setae; protibial anterior margin with 6 spines. Abdomen: Aedeagus as in figure 27.

Variation. One specimen examined.

Natural History. The Holotype was collected during June at 216 m.

Distribution. From the Reserva Ethnica Waorani and the environs of the Yasuni National Forest of eastern Ecuador.

Etymology. The specific epithet demagna is a Latin compound name derived from de (= very) and magnus (= large). I refer to the extraordinarily large last antennomere.

Taxonomic Notes. Among the Amboakis species keyed in Opitz (2006: 119) A. demagna is closely related to the Brazilian A. capitata and A. antegalba . The following changes are made to the aforementioned key:

19'. Last antennomere not more than twice as long as penultimate antennomere ........................ 21

20(19). With narrow yellow line projecting backwards from elytral humeral margin ( Ecuador: Orellana)

......................................................................................... Amboakis demagna Opitz , n. sp. – Without narrow yellow line projecting backwards from elytral humeral margin ( Brazil: Mato

Grosso; Nova Teutonia; Paraná) ..................................................... Amboakis prolata Opitz

Figures

Figure 1-15. Antennae. 1) Amboakis ampla. 2) Amboakis capitata. 3) A. stenosis. 4) A. waodani. 5) Madoniella aspera. 6) M. divida. 7) Plocamocera quadrula. 8) P. clinata. 9) Amboakis antegalba. 10) A. diffusa. 11) Madoniella spilota. 12) Amboakis demagna. 13) Epiphloeus erwini. 14) Madoniella darlingtoni. 15) Plocamocera lena.

Figure 25-38. Aedeagi. 25) Amboakis ampla. 26) A. antegalba. 27) Amboakis demagna. 28) Amboakis capitata. 29) A. diffusa. 30) A. waodani. 31) Epiphloeus erwini. 32) Epiphloeus pulcherrimus. 33) Madoniella divida. 34) Plocamocera lena. 35) Pyticeroides latisentis. 36) P. moraguesi. 37) P. turbosiris. 38) P. ustulatis.

Figure 47-58. Habiti. 47) Amboakis ampla. 48) A. antegalba. 49) A. diffusa. 50) Amboakis demagna. 51) A. waodani. 52) Epiphloeus erwini. 53) Madoniella aspera. 54) Madoniella darlingtoni. 55). Madoniella divida. 56) Madoniella spilota. 57) Plocamocera clinata. 58) P. lena.