Heresiarchini ASHMEAD

A. M, 2009, Illustrated key to the tribes of subfamilia Ichneumoninae and genera of the tribe Platylabini of world fauna (Hymenoptera, Ichneumonidae), Linzer biologische Beiträge 41 (2), pp. 1317-1608 : 1386-1387

Tribe Heresiarchini ASHMEAD

Heresiarchini ASHMEAD 1900 - Proc. U.S. Nat. Mus. 23 (1206): 1-220.

Type genus: Heresiarhes WESMAEL.

Heresiarchina HEINRICH 1970 - Entom. Tidskr. 91: 68-69.

Heresiarchini : PERKINS 1959 - Roy. Ent. Soc. London 7, 2 (aii): 24-25, 45.

Heresiarchini : RASNITSYN 1981 - Opr. Faun. SSSR 3 (3): 508.

D i s t r i b u t i o n: Palaearctic (1 species) and Oriental Region.

I n t r o d u c t i o n:

The tribe is characterized by combination of two characters: 1 – sickle-shaped mandibles with long upper tooth and atrophied lower one, bent to internal margin of a mandible and 2 – sloped propodeum with peculiarity reduced fields.

The similar morphology of mandibles is peculiar to the representatives of Joppocryptini and to a number of genera of subtribe Cratichneumonina of tribe Ichneumonini (for example Rhadinodonta SZEPLIGETI and Hymenura TOWNES (= Neopyga HEINRICH).

Propodeum of Heresiarchini from lateral uniformly slanted from base to apex thus, that third pleural area (area posteroexterna) extremely sharply shortened, hereupon the apices of areae dentiparae especially close approach to the hind coxae. The fields of propodeum are reduced: area dentipara and area posteroexterna completely merged, fields of horizontal part, area superomedia, area basalis and area superoexterna are merged, forming single polished surface. By morphology of propodeum, as per opinion of G. Heinrich one of the most significant segment for Ichneumoninae , Heresiarchini are closest to tribe Protichneumonini (African subtribe Apatetorina ), even if are not a part of it.

Oxypygous abdomen with straight from lateral first tergite and deep gastrocoeli with small, but distinct thyridia are also typical to the representatives of the tribe.

M o r p h o l o g y:

F l a g e l l u m: Of females slender, long, bristle-shaped, slightly widened and flattened ventrally. Flagellum of males with tyloides.

H e a d: Temples short, behind eyes from above sharply roundly narrowed; mandibles sickle-shaped, slender, lower tooth far moved inside and practically not visible from front; clypeus flat, slightly impressed relatively face.

T h o r a x: Mesonotum moderately convex, of equal length and breadth; notauli completely absent; sternauli practically absent, only in form of broad slight impressions at base; scutellum practically not elevated, flat, only just carinated. Hind margin of metanotum with projections. Propodeum from lateral uniformly sloping from base to apex, without dividing on horizontal and vertical parts; fields reduced: area dentipara and area posteroexterna entirely merged; area superomedia merged with area basalis and area superoexterna, forming single solid surface; third pleural area (apical pleural area) extremely sharply shortened, as result apices of areae dentiparae especially close approach to hind coxae; coxal and metapleural carinae sharp; spiracles long, slit-shaped. Central part of propodeum up to apical transverse carina not sculptured, polished.

L e g s: Thin and slender; claws long, smooth, slightly curved; hind coxae of female without scopa.

W i n g s: Areolet pentagonal.

A b d o m e n Abdomen from above long, fusiform, seventh tergite sharpened, long, ovipositor distinctly protrude behind apex; abdomen oxypygous. First tergite from lateral practically straight; from above with very narrow petiolus at base; from lateral smooth, without traces of lateral carinae and any sculpture; gastrocoeli deep, triangle, but small; thyridia distinct.

C o l o r a t i o n: Abdomen with white spots at corners of front tergites and developed bands and spots on apical tergites.