Spinosciacharis acheron, Jałoszyński, Paweł, 2014

Jałoszyński, Paweł, 2014, ' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae), Zootaxa 3828 (1), pp. 1-76 : 49-50

publication ID	https://doi.org/ 10.11646/zootaxa.3828.1.1
publication LSID	lsid:zoobank.org:pub:B2FEAE60-7E51-45FA-A38F-930A084A5AAA
DOI	https://doi.org/10.5281/zenodo.6141084
persistent identifier	https://treatment.plazi.org/id/03F68791-FFE7-1839-8FFE-387003A3FD59
treatment provided by	Plazi
scientific name	Spinosciacharis acheron
status	sp. nov.

Treatment

Spinosciacharis acheron View in CoL sp. n.

( Figs. 97–111 View FIGURES 97 – 101 View FIGURES 102 – 104 View FIGURES 105 – 108 View FIGURES 109 – 111 , 116 View FIGURES 112 – 116 , 120 View FIGURES 117 – 120 , 127–128 View FIGURES 121 – 128 , 196 View FIGURES 193 – 198 d)

Type material studied. Holotype: AUSTRALIA: ♂: four labels: " AUSTRALIA / Victoria / Acheron Gap / 750m nr. / Warburton / 28-30 Apr. 78 / S. & J. Peck" [white, printed], "Berlesate frass / under Nothophagus / logs" [white, printed], "Aust. Nat. / Ins. Coll." [green, printed], " Spinosciacharis / acheron m. / det. P. Jałoszyński, 2014 / HOLOTYPUS " [red, printed] ( ANIC). Paratypes (2 ♂♂, 3 ♀♀): same data as for the holotype, all with standard yellow " PARATYPUS " label ( ANIC, cPJ).

Diagnosis. Male and female: BL 1.78–1.98 mm; elytra oval; EI 1.46–1.56. Male: metafemur with distinctly recurved and broadly rounded ventral projection, with distinct distal ventral lobe.

Description. Body of male ( Fig. 116 View FIGURES 112 – 116 ) flattened, elongate and slender, with long appendages, BL 1.90–1.98 mm (mean 1.93 mm); cuticle glossy, body light brown, with slightly lighter appendages, vestiture yellowish.

Head ( Fig. 116 View FIGURES 112 – 116 ) rounded, broadest just behind eyes, HL 0.38–0.40 mm (mean 0.38 mm), HW 0.40–0.43 mm (mean 0.41 mm); tempora nearly three times as long as eyes, strongly curved posteromesally; vertex evenly convex; frons confluent with vertex, subtrapezoidal, convex. Eyes small and weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum inconspicuous, fine and sparse, not larger than those on pronotum; setae on frons and median part of vertex short, sparse and suberect, tempora and sides of vertex with moderately dense bristles directed posterolaterally. Antennae ( Fig. 116 View FIGURES 112 – 116 ) slender, AnL 0.90–0.93 mm (mean 0.92 mm); antennomeres I–VII distinctly elongate, VIII about as long as broad, IX-X distinctly transverse, XI much shorter than IX–X together, 1.6x as long as broad, pointed.

Pronotum ( Fig. 116 View FIGURES 112 – 116 ) in dorsal view elongate, broadest near anterior third, PL 0.50 mm, PW 0.40–0.45 mm (mean 0.42 mm); anterior and lateral margins rounded; posterior margin indistinctly bisinuate; base of pronotum with distinct but short sublateral carinae accompanied mesally by shallow impressions, median part of pronotal base with two small but deep pits separated by narrow carina twice as long as pits and extending beyond pits only posteriorly, up to posterior pronotal margin. Punctures on pronotal disc fine and inconspicuous, similar to those on frons and vertex; setae sparse, thin, short and suberect, only sides of pronotum with moderately dense bristles and in anterior half with several extremely long, erect setae directed laterally.

Elytra ( Fig. 116 View FIGURES 112 – 116 ) slightly rhomboidal, broadest distinctly anterior to middle, EL 1.03–1.08 mm, EW 0.68–0.73 mm, EI 1.48–1.56; basal impressions short but deep; circumsutural area in anterior fourth slightly more convex than surrounding surface and posteriorly accompanied by distinct, shallow and short impression; elytral apices rounded together. Punctures on elytral disc fine but slightly more distinct and denser than those on pronotum; setae short, sparse and suberect. Hind wings well developed, about twice as long as elytra.

Legs ( Figs. 116 View FIGURES 112 – 116 , 120 View FIGURES 117 – 120 ) long and slender, metafemur ( Fig. 120 View FIGURES 117 – 120 ) with long and distinctly recurved ventral projection slightly narrowing toward broadly rounded apex, with distinct ventral distal lobe.

Aedeagus ( Figs. 127–128 View FIGURES 121 – 128 ) relatively stout, AeL 0.48 mm, in ventral view sides of median lobe rounded, apical margin subtriangular and broadly rounded; internal armature lightly sclerotized, composed of elongate paired sclerites; parameres not broadening distally and their apices only slightly exceeding apex of median lobe.

Female. Similar to male, differs in unmodified metafemora; BL 1.78–1.95 mm (mean 1.87 mm); HL 0.38 mm, HW 0.38–0.43 mm (mean 0.40 mm), AnL 0.83–0.90 mm (mean 0.86 mm); PL 0.43–0.50 mm (mean 0.47 mm), PW 0.40–0.43 mm (mean 0.42 mm); EL 0.98–1.08 mm (mean 1.03 mm), EW 0.63–0.70 mm (mean 0.68 mm), EI 1.46–1.56 (mean 1.52).

Etymology. Locotypical, after the Acheron Gap.

Distribution ( Fig. 196 View FIGURES 193 – 198 d). South-eastern Australia: Victoria.

Remarks. This species is most similar to Spinosciacharis clydensis ; see Remarks section under the latter species.