Tipula (Sivatipula) podenasi, Men, 2020

Men, Qiulei, 2020, Three new species of the subgenus Tipula (Sivatipula) from Yunnan and Tibet, China (Diptera: Tipulidae), with an updated key for Chinese species, Zoological Systematics 45 (1), pp. 59-68 : 65-68

Tipula (Sivatipula) podenasi sp. nov. ( Figs 30–43 )

Diagnosis. Body brownish-yellow in general; thorax brownish-yellow with three light brown stripes on prescutum, the median one divided by a narrow yellow vitta; antenna longer than the body length; wing brownish-yellow with a light brown stigma; abdomen brownish-yellow with lateral stripes black, segments five to seven wholly black, hypopygium black, distinctly enlarged; tergite nine with a pair of horn-shaped processes at posterior margin; outer gonostylus huge, boomerangshaped.

Material examined. Holotype male, China, Yunnan, Gongshan, Bingzhongluo, Gongdangshenshan , 28°01'N, 98°37'E, 12 Jun 2019, leg. Qiulei Men. Paratype. 1 male, China, Yunnan, Gongshan, Shuanglawa , 27°46'N, 98°37'E, 13 Jun 2019, leg. Qiulei Men; 1 male, same data to holotype .

Description. Adult length (male, n = 3): body 16.1–16.3mm (not including antenna); wing 15.0– 15.2mm; antenna 17.8– 18.0 mm.

Head. Brownish-yellow with rostrum brown in lateral sides. Nasus brownish-yellow densely covered with black setae ( Fig. 30 ). Vertex lacking of marking ( Fig. 30 ). Head including vertex and rostrum densely covered with black setae. Antenna 12-segmented, significantly longer than body; scape and pedicel brownish-yellow, first flagellomeres brownish-yellow at base and brown apically, rest flagellomeres brown, each flagellomere subequal in length, cylindrical, with black verticils. Palpus brown.

Thorax. Pronotum brownish-yellow ( Fig. 31 ). Prescutum brownish-yellow with three light brown stripes, the median one divided by a narrow yellow vitta ( Fig. 32 ). Scutum brownish-yellow with two brown triangular marks which are not connected to each other, scutellum and postnotum brownish-yellow ( Fig. 32 ). Pleuron entirely brownish-yellow ( Fig. 31 ). Legs very slender, coxae and trochanters brownish-yellow, femora brownish-yellow basally and gradually changed into light brown, dark brown at apex, tibiae and tarsi dark brown. Halter with stem yellow, knob darker. Wing brownish-yellow, cell sc not darker than ground color, stigma light brown ( Fig. 33 ). Rs distinctly longer than R 3, petiole of cell m 1 distinctly shorter than m-m, 1/4 the length of discal cell ( Fig. 33 ).

Abdomen. Segments one to five brownish-yellow with black lateral stripes, segments six to eight wholly black ( Fig. 34 ). Hypopygium black, distinctly enlarged ( Fig. 35 ). Tergite nine with a pair of horn-shaped processes, between them with a small angle slightly protruded medially, with a pair of arms on ventral side which expand apically and equipped with many small black teeth ( Figs 36–38, 41). Outer gonostylus huge, narrowed, curved, boomerang-shaped, more than two times longer than the length of inner gonostylus ( Figs 36, 39). Inner gonostylus broad at base, gradually narrowed to apex, with a small angular lobe on the lateral margin, an elongated obtuse lobe generated from the base, a small lobe arising from its mesal region of inner surface ( Figs 39–40). Sternite nine with a pair of short, finger-shaped processes densely covered with small setae ( Fig. 38).

Semen pump with compressor apodeme V-shaped, each arm broad ( Fig. 42). Posterior immovable apodeme being single median arm which is curved dorsally ( Fig. 43). Anterior immovable apodeme flattened ( Fig. 43). Aedeagus relatively thick, more than 3.5 times longer than semen pump ( Fig. 43).

Distribution. China (Yunnan).

Remarks. The new species is most similar to T. (S.) bhishma by similar structure of hypopygium. It can be easily distinguished from the latter by outer gonostylus which is broad basally and narrowed to apex. T. (S.) bhishma has its outer gonostylus broad at apex. The new species is also distinguished by the horn-shaped process on tergite nine, which is distinctly narrower than that of T. (S.) bhishma .

Etymology. The new species is named in honor of famous crane flies researcher, Sigitas Podenas (Nature Research Centre, Vilnius, Lithuania).

Figures

Figures 30–35. Tipula (Sivatipula) podenasi sp. nov. 30. Head, dorsal view. 31. Thorax, lateral view. 32. Thorax, dorsal view. 33. Wing. 34. Abdomen and hypopygium, lateral view. 35. Hypopygium, lateral view.