Eusphalerum uncinatum Zanetti, 2014

Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), Insecta Mundi 2014 (379), pp. 1-80 : 57-58

Eusphalerum uncinatum Zanetti n. sp.

Material examined (135 specimens)

Holotype m 5 paratypes mm 7 ff Oregon Clackamas Co Camp Crk. (3.5 mi. SE) 2300-2400' Rhododendron 27.06.1974 leg. A. & D. Smetana (CNC)

Other paratypes. CANADA. British Columbia 18 mm 18 ff Manning Pr. Pk Rhododendron Flats Rhododendron flowers 21.06.1988 leg. J.M. Campbell (CNC); 6 mm 5 ff Manning Prov. Pk., Rhododendron Trail sweep Rhododendron flowers 20.06.1988 leg. L. LeSage (CNC); 2 mm Mt. Revelstoke 6000 25.07.1972 leg. G.J. Spencer (CNC); 1 m Silver Star Prov. Pk. 6200 8.08.1973 leg. R.H. Parry (CNC). USA. California 10 m 33 ff El Dorado Co Fallen Leaf 6500 1/ 29.07.1935 leg. F.E. Blaisdell (CNC); 1 m El. Dorado Co Fallen Leaf 21.06.1915 leg. R.Hopping (CNC); 3 mm 2 ff Fresno Co Huntington Lake swept 3.08.1962 leg. C. D. Johnson (FMNH); 6 mm 3 ff Fresno Co Sierra NF John Muir Tr 37.2583 oN 118.8654 oW 20.08.2006 leg. Caterino (SBMNH); 1 m 1 f Placer Co Lake Tahoe, Tahoe Pines 6200 10.08.1969 leg. A. Smetana (CNC); 2 mm Tehama Co Wilson Lake 5300 27.06.1966 leg. C.W. OBrien (CNC); 2 f 1 m Lake Tahoe 23.06.1925 leg. E.H.Nast (CNC); 2 mm Fresno Co Huntington Lake 3.08.1962 leg. C.D. Johnson (FMNH); Oregon 1 m Clackamas Co jct US26 Ore 35 (1.3 m E) 3900 on flowers Lupinus 12.07.1975 leg. A. Newton M. Thayer (FMNH); 1 m 1 f Douglas Co Lamola Lk. (3.5 mi. S), Umpqua Nat. For. 4400' currant, Pinus contorta , rotted wood 22.06.1972 leg. E.H. Benedict (CNC); Washington 2 mm 2 ff Chelan Co Leavenworth (10 km NW) 30.06.1984 leg. R. Baranowski (LMZ).

Measurements. head length: 0.31-035; head width: 0.53-0.62; pronotal length: 0.46-0.55; pronotal width: 0.72-0.83; elytral length: 0.98-1.14; elytral width: 0.99-1.18; length (clypeus to apex of elytra): 1.66-2.23; total length: 2.1-3.1.

Etymology. The adjective uncinatum means hooked in Latin. It refers to the form of median lobe of the aedeagus in lateral view.

Description. Habitus as in Fig. 149. Head and pronotum reddish yellow; neck sometimes somewhat darkened in male; elytra yellowish;, abdomen brown, somewhat paler at apex (male) or entirely yellowish (female); prosternum yellowish; metasternum brown, sometimes yellowish in female; legs yellowish; antennae yellowish, more or less darkened from antennomere 6, often entirely yellowish.

Head with prominent eyes; postocular carina well marked; temples short, convergent caudad. Medial margin of eyes with microsculpture forming longitudinal wrinkles; postantennal depressions deep, confluent with tentorial pits, forming 2 longitudinal depressions in front of ocelli; punctation superficial and sparse; ground with microsculpture isodiametric in middle and on neck, longitudinal in depressions in front of ocelli; neck wide, not separated from head. Antennae robust, antennomere 1 and 2 ovoid, 3 elongate, twice as long as wide, 4, 5, and 6 subquadrate, 9-10 transverse, 10 almost twice as wide as long, 11 twice as long as wide, cylindrical in the proximal part, conical at apex.

Pronotum convex, often with a small impression medially near anterior margin, transverse (ratio width/length = 1.4-1.6), widest in middle, anterior margin slightly narrower than posterior, lateral margins usually rounded, sometimes almost rectilinear, posterior angles scarcely obtuse and well marked, punctation sparse and rather superficial, ground with isodiametric microsculpture, pubescence extremely short, scarcely visible, depressions near posterior angles rather wide.

Elytra scarcely elongate (ratio length from scutellum to apex / combined width of elytra = 1.0), truncate at apex in both sexes, moderately widened toward apex, punctation stronger than on pronotum, irregular and rather confluent, ground glossy, pubescence extremely short and scarcely visible.

Abdomen rather glossy, microsculpture superficial with decumbent pubescence.

Tibiae straight in both sexes, not modified, tarsomere 5 of posterior tarsi slightly shorter than 1-4 together.

Aedeagus as in Fig. 150, hooked near apex in lateral view, without large copulatory sclerites.

Accessory sclerites of female as in Fig. 151, spermatheca as in Fig. 152.

Comparative notes. Eusphalerum uncinatum is similar to E. pothos , from which the absence of a defined median groove on pronotum is somewhat distinctive. Eusphalerum newtoni and E. parvispiculum are similar, too, and only the shape of the aedeagus, especially the presence of a hook near the apex, visible in lateral view, is distinctive for E. uncinatum .

Distribution. CANADA: British Columbia; UNITED STATES: California, Oregon, Washington (Map 9).